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From: "Helmut Zibrowius" <hzibrowi@com.univ-mrs.fr>
Date: Fri, 1 Dec 95 12:00:07 MST
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To: annelida@net.bio.net
Subject: Sabella spallanzanii from Java?

CHAETZONE no. 9 edited by G. Read contains some information on S. 
spallanzanii, part of which which is quoted here: 
-------------------------------------

* - SABELLA SPALLANZANII DISTRIBUTION
    -------------------------------
    From Phyllis Knight-Jones <pknight-jones@bryngwyn.demon.co.uk>

  As readers of Chaetozone
  will know there has been great interest in the discovery of a vast
  population of this species in Geelong harbour near Melbourne since Carey
  and Watson's report (Carey J.M. and Watson J.E., 1992. Benthos of the
  muddy bottom habitat of the Geelong arm of Port Phillip Bay, Victoria,
  Australia. _The Victorian Naturalist_ pp. 196-202.) I examined the Siboga
  material from a reef off Nusa-Laut Island, Indonesia (46m depth), but it
  is not _Sabella spallanzanii_ ('Spirographis Spallanzanii'). It is in
  fact _Bispira tricyclia_ (Schmarda), the only unispiral _Bispira_ in the
  genus (as amended in Perkins and Knight-Jones, 1991). The two species
  were wrongly synonymised by Fauvel (eg. 1953).

  After very interesting studies on the Amsterdam collections (various
  genera), Harry ten Hove accompanied us to Leiden and had the foresight to
  search for unregistered (unidentified) sabellid collections, while I
  browsed through official holdings. Knowing of my interests in both
  Indonesia and _Sabellastarte_, he selected a jar of very large sabellids
  labelled 'Kuhl, Java'. Examination proved this to be six specimens of
  _Sabella spallanzanii_. Jacob van der Land informed us that Dr Kuhl, a
  young short-lived medic, collected these in 1820, but no further details
  of location are available. Kuhl was a mainly terrestrial naturalist,
  perhaps without naval support, so one might speculate that the material
  was gathered in a convenient harbour, perhaps Batavia, now known as
  Djakarta.

  Now this Indonesian material is identified, 165 years after it was
  collected, we have reliable records of _Sabella spallanzanii_ from the
  Mediterranean, NW France, Azores, Rio de Janeiro, Java and Australia. The
  oldest records are from the Mediterranean and Java, the next oldest
  probably Hanson's (1882) descriptions from Rio under 4 different names,
  all synonyms with _spallanzanii_. It is interesting to note that these
  locations coincide with the old sailing route to Asia, though a notable
  absence is the revictualling stop at Cape Town. There, however, the
  original harbours are long since under concrete. -- Phyllis Knight-Jones
------------------------------------
As I already informed Phyllis, I have a comment concerning the KUHL Java 
1820 record.

   I would be less confident in this record. Have come across many 
errors concerning old material, and including the Challenger! But I am 
confident in Jacob van der Land's detective and archeologist work: that 
Kuhl was a medic who traveled to Java around 1820.

   Take the following case which, I presume, may throw some doubt on 
the Javanese origin of those old Leiden S. spallanzanii:

   My detailed investigations on the colonial scleractinian coral 
Sclerhelia hirtella (Pallas, 1766) suggest that this species is a true 
endemic of St. Helena, South Atlantic.
(Zibrowius H., 1974. Redescription of Sclerhelia hirtella from Saint Helena,
South Atlantic, and remarks on Indo-Pacific species erroneously referred to 
the same genus (Scleractinia). Journal of natural history, 8 (5): 563-575. -
- Further information to be included into a more comprehensive coral fauna 
of St. Helena, in preparation.)
   St. Helena is known as the British base where the Corsican/French 
military dictator Napoleon had been exiled to. Before the opening of the 
Suez Canal the isalnd had some importance as a revictualling stop 
on the way to and from India.
   Pallas (1766), the author of Sclerhelia hirtella (first illustration by 
Ellis & Solander, 1786) presented the species as "very rarely received from 
the Indian Ocean". INDIAN OCEAN as the origin of the species has been 
reiterated several times until 185O. Only in 1857 Milne Edwards & Haime 
have been able to indicate St. Helena as the true origine of that species, 
on the basis of more precisely recorded material. In fact, at St. Helena 
the species is common and lives from a few meters depth to deeper water, 
thus accessible to anchor lines and fishing gear even in former times.
No surprise that occasionally it found its way into natural history 
cabenets in Euriope.

   Back to S. spallanzanii and having in mind the coral story above:   
   Kuhl may well have got his sabellid worms somewhere in western Europe on 
his way BACK from Java.

   Or perhaps from the hull of the ship that took him to Java???

   See the following case: A sample of Hydroides elegans, the now worldwide 
fouling serpulid in tropical and warm-temperate waters (harbour communities 
and ship hulls, has well been brought back from the remote St. Paul and 
Amsterdam islands, southern Indian Ocean (halfway between South Africa and 
Australia): not from (unexisting) harbours there, but from the cooling 
water intake sieve of a French vessel fishing spiny lobsters around those 
islands.

   Some further comments or similar observations?

  ----------------------------------- 
  Helmut ZIBROWIUS
  (Centre d'Oceanologie de Marseille)
  Station Marine d'Endoume
  Rue Batterie des Lions
  13007 Marseille / France
  TEL: (intern. - 33) 9 1 0 4 1 6 2 4
  FAX: (intern. - 33) 9 1 0 4 1 6 3 5
  E-MAIL: <hzibrowi@com.univ-mrs.fr>
  -----------------------------------

From BIOSCI-REQUEST  Mon Dec  4 07:47:17 1995
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From: Geoffrey Read <Geoffrey.Read@actrix.gen.nz>
Message-Id: <199512040221.PAA18783@atlantis.actrix.gen.nz>
Subject: Re: Sabella spallanzanii from Java?
To: annelida@net.bio.net (ANNELIDA list)
Date: Mon, 4 Dec 1995 15:21:34 +1300 (NZDT)
In-Reply-To: <9295.hzibrowi@[139.124.16.1]_POPMail/PC_3.2.2> from "Helmut Zibrowius" at Dec 1, 95 12:00:07 pm
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Helmut Zibrowius writes:-

[quotes from Phyllis Knight-Jones] ...
>   "Indonesia and _Sabellastarte_, he selected a jar of very large sabellids
>   labelled 'Kuhl, Java'. Examination proved this to be six specimens of
>   _Sabella spallanzanii_.
    [...]
>   Now this Indonesian material is identified, 165 years after it was
>   collected, we have reliable records of _Sabella spallanzanii_ from the
>   Mediterranean, NW France, Azores, Rio de Janeiro, Java and Australia...."

[then comments]
>    I would be less confident in this record. Have come across many 
> errors concerning old material, and including the Challenger! But I am 
> confident in Jacob van der Land's detective and archeologist work: that 
> Kuhl was a medic who traveled to Java around 1820. ...

I enjoyed Phyllis's latest detective story. As an outsider it does seem that
reliable is perhaps too strong a word for the record (Do we have a volunteer
to check it out in Jakarta?). Short of finding Kuhl's diary ("today during
careenage at X collected strange growths") or supporting evidence from other
Kuhl stuff in the museum we'll never know will we? What is the suggested
scenario here anyway? That _S. spallanzanii_ was exported from Europe, or the
reverse, or neither? Reminds me of the similarly uncertain _Ficopomatus
enigmaticus_ situation, to which no doubt someone will apply the appropriate
modern genetic tools eventually. 


Geoff 
-- 
   Geoff Read <gread@actrix.gen.nz>
   Annelida resources =>  http://www.actrix.gen.nz/users/chaeto/index.html

From BIOSCI-REQUEST  Mon Dec  4 09:13:00 1995
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From: Geoffrey Read <Geoffrey.Read@actrix.gen.nz>
Message-Id: <199512040524.SAA14126@atlantis.actrix.gen.nz>
Subject: The ANNELIDA archive (useful administrivia)
To: annelida@net.bio.net (ANNELIDA list)
Date: Mon, 4 Dec 1995 18:24:49 +1300 (NZDT)
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Hello Annelidans,
 
If you need to review the messages posted since
ANNELIDA began two weeks ago the WWW address is:-
 
gopher://gopher.bio.net:70/11/ANNELIDA
 
All messages are automatically placed there via Bionet's archiving system. 
Each month will be in a separate 'folder' (9511, 9512). When you click on 
the folder you see the list of subject headers and can open each message 
individually. Very neat!
 
By the way, after posting a message to annelida@net.bio.net it takes awhile 
before it can be sent to everyone. Don't be surprised if there is a delay 
of an hour or so before you receive back your own copy (whereas you 
will get a very quick acknowledgement to unsubscribe/subscribe messages to 
biosci-server@net.bio.net).
 
CHAETOZONE NINE polychaete newsletter was mailed on 1 December. Brant 
Mitchell, Robin Hensley, David Bone Torroja might like to get in touch if 
still listening out there as your copies 'bounced'. Anyone else interested 
can also make contact or look on the gopher sites, gopher.ucsc.edu, 
muse.bio.cornell.edu, in a few days time.

Cheers,

Geoff
-- 
   Geoff Read <gread@actrix.gen.nz>
   Annelida resources =>  http://www.actrix.gen.nz/users/chaeto/index.html

From BIOSCI-REQUEST  Mon Dec  4 15:34:11 1995
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unsubscribe annelida


From BIOSCI-REQUEST  Mon Dec  4 18:19:23 1995
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From: Geoffrey Read <Geoffrey.Read@actrix.gen.nz>
Message-Id: <199512050217.PAA23746@atlantis.actrix.gen.nz>
Subject: Re: Sabella spallanzanii from Java? (fwd)
To: annelida@net.bio.net (ANNELIDA list)
Date: Tue, 5 Dec 1995 15:17:39 +1300 (NZDT)
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===================== Begin forwarded message ==========================

Date: Mon, 04 Dec 1995 12:43:22 -0500
From: Kristian Fauchald <mnh.fauchald@ic.si.edu>
To: gread@actrix.gen.nz
Subject:  Re: Sabella spallanzanii from Java? -Reply

Several hundred years had passed between the "discoveries" of the
various non-European countries and the time we started to become
aware of the potential for transportation on ship-bottoms and so forth. 
There is something a bit peculiar about the whole debate about
transportation: 1. Rafting on pieces of wood or on ship-bottoms is
different in kind apparently from the rafting on pieces of continent.  Time
scale is different, but in principle rafting is rafting whether it is slow or
fast.  2.  The fact that the earliest description might be from Europe or
North America does not guarantee that this is the "origin" of the taxon: 
All it says is that the taxon was first noticed where the density of
taxonomists where the highest. 

Kristian Fauchald
NHB MRC 163
Smithsonian Institution
Washington, DC 20560 USA
Tel. (202)357-4757; FAX (202)357-3043
e-mail MNH.FAUCHALD@IC.SI.EDU


From BIOSCI-REQUEST  Mon Dec  4 21:10:27 1995
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From: jablake@ix.netcom.com (JAMES BLAKE )
Subject: Re, type localities and distributions of species
To: annelida@net.bio.net

Dear annelid enthusiasts, 

The recent exchange of information about Sabella spallanzanii opens an 
interesting area of discussion, namely that of the importance of type 
localities for species that have become widely distributed due to man's 
activities.  The question might be raised as to what importance is the 
type locality, other than than the source of the original material, for 
a species that has been widely dispersed in ballast water, mud 
adhereing to seed oysters, or attached to a growth on a ship's hull? I 
suppose I could ask the question--Who cares about the type locality?  
Of what importance is such information?  

There are several examples where knowledge of the type locality has 
actually become misleading in understanding the origins of species or 
species groups.  In San Francisco Bay, there are essentially no 
endemics; virtually all of the fauna has been introduced in some 
manner.  There are quite a few examples of invertebrates that have 
species epithets of "californiensis" or "franciscanus" that were 
introduced into San Francisco Bay from some exotic locality.  In some 
instances, the origin of such species is still not known.  Jim Carlton 
has pointed out that the homeland of the isopod, Synidotea laticauda, 
originally described from SF Bay, is still unknown.  In Tomales Bay, 
both of the spionids, Pseudopolydora kempi (Southern) and P. 
paucibranchiata (Okuda) were introduced sometime in the late 1960's or 
early 1970's via seed spat of the Pacific Oyster imported from Japan.  
Seed oysters contain considerable mud packed between shells and when 
laid out on tidal flats there is nothing to prevent small worms and 
other invertebrates from staking out their own territories.  I suspect 
that quite few of the nereids, syllids, and other polychaetes common in 
Tomales Bay and other estuarine environments of California have 
probably been introduced (and are still being introduced) into these 
waters.  Any species (adults or larvae of same) that is euryhaline 
should have some chance of surviving in ballast water shipments as Jim 
Carlton has so convincingly demonstrated in his experiements. 

So, I ask the question as to what importance is the type locality in 
understanding a species, if it has been moved around from place to 
place and the actual area of the world where it originated is somewhere 
quite different?  

I recall a conversation I once had with a colleague who refused to 
accept an identification I had made because the type locality was 
somewhere far distant from where the specimens being identified were 
collected.  When I asked the question as to what the type locality had 
to do with the distribution of a species, I was met with a very puzzled 
look.  The answer was something like, "Why, that is where it is from."  
I think you see the point, namely that type localities are only the 
places where the animals were first collected, not necessarily where 
they are most common or where they originated.  

Does anyone out there have any comment on this issue or examples of 
species that have obviously been moved around by ships or other means?  



Jim Blake
ENSR
89 Water Street
Woods Hole, MA 02543
(jablake@ix.netcom.com)



From BIOSCI-REQUEST  Tue Dec  5 00:49:56 1995
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From: Geoffrey Read <Geoffrey.Read@actrix.gen.nz>
Message-Id: <199512050848.VAA28999@atlantis.actrix.gen.nz>
Subject: Errant earthworms (reply re Sabella etc) (fwd)
To: annelida@net.bio.net (ANNELIDA list)
Date: Tue, 5 Dec 1995 21:48:04 +1300 (NZDT)
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Help! I'm trapped in a forwarding loop!

Intended for <annelida@net.bio.net>

For better or worse I've changed the subject header.  Please reply to Sam, or
preferably continue this discussion on the Annelida list. -- GBR

===================== Begin forwarded message ===========================
Sam James writes:-
Date: Tue, 5 Dec 95 00:47:53 CST
From: "Sam James" <sjames@mum.edu>
Subject: Re: Sabella spallanzanii from Java? (fwd)

>Date: Mon, 04 Dec 1995 12:43:22 -0500
>From: Kristian Fauchald <mnh.fauchald@ic.si.edu>
>To: gread@actrix.gen.nz
>Subject:  Re: Sabella spallanzanii from Java? -Reply
>
>Several hundred years had passed between the "discoveries" of the
>various non-European countries and the time we started to become
>aware of the potential for transportation on ship-bottoms and so forth. 
>There is something a bit peculiar about the whole debate about
>transportation: 1. Rafting on pieces of wood or on ship-bottoms is
>different in kind apparently from the rafting on pieces of continent.  Time
>scale is different, but in principle rafting is rafting whether it is slow or
>fast.  2.  The fact that the earliest description might be from Europe or
>North America does not guarantee that this is the "origin" of the taxon: 
>All it says is that the taxon was first noticed where the density of
>taxonomists where the highest. 
>
>Kristian Fauchald

Hear, Hear! The situation is paralleled by earthworms, which 
raft on continents and islands only (except for the very few euyhaline 
species), as far as is known . Some have achieved wide distributions 
cruising about _inside_ ships and airplanes.  There are many cases where 
species were first described from material collected well outside the 
natural ranges of the species or their close relatives.  For example: 
Drawida bahamensis, a native of India first found halfway around the world 
from its origin.
    The only situation of which I am aware that has caused any debate 
about natural ranges involves the Lumbricidae, a Holarctic but largely 
Palearctic family with a couple dozen species common to both continents.  
Now it is generally believed that these species owe their residence in 
North America to human activity.  It helps that they are also quite common 
in south temperate and montane tropical areas.
   Are most polychaetes confined to fairly limited areas, with a few 
"naturally" widely distributed species?

~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~
Sam James    Dept. of Biology
FM 1056   Maharishi Univ. of Management
Fairfield, Iowa USA 52557
515-472-1146  fax:472-1167

Ecology and Systematics of Earthworms
~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~

From BIOSCI-REQUEST  Tue Dec  5 06:37:13 1995
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*********************************************
Dr. Stephen R. Moulton II
Biologist
United States Geological Survey
National Water Quality Laboratory
Biological Quality Assurance Unit
5293 Ward Road, MS 426
Arvada, CO  80002

office: (303) 467-8171
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From BIOSCI-REQUEST  Tue Dec  5 10:03:05 1995
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From BIOSCI-REQUEST  Tue Dec  5 10:03:40 1995
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Date: Tue, 5 Dec 1995 10:59:03 +0000
From: Dani Martinez <dani@azathoth.ceab.es>
Sender: Dani Martinez <dani@azathoth.ceab.es>
Reply-To: Dani Martinez <dani@azathoth.ceab.es>
Subject: Type localities and dispersion of species
To: annelida@net.bio.net
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Dear Anelidans,

Talking about the importance of type localities,
Jim Blake writes:-
> Does anyone out there have any comment on this issue or examples of
> species that have obviously been moved around by ships or other means?

=09Concerning type localities, I had the opportunity to describe a=20
new capitellid genus from a brackish, shallow-water, enclosed Bay of the=20
Ebro'=D5s Delta in the Iberian peninsula which I named" =D2Pseudomass tu
deltaicus"=D3. It was very abundant there (about 2000 individuals per squar=
e=20
meter). As Mediterranean waters have been (and are being) largely=20
explored, I thought the species was endemic. I mentioned this possibility=
=20
in the original description, and it was the main reason when I decided=20
this =D2restrictive=D3 name. A few years latter I was really surprised when=
=20
some colleagues start to mention this species from areas different (and=20
far) of the type locality. Moreover, most of these citations were from=20
open bays (although the influence of continental waters were always=20
present).=20
=09I do not know whether this could be a case of recent migration,=20
previous misleading identifications, hazard of looking now at the right=20
zones, or whatever. The main point is that the name of the species,=20
directly linked to the type locality, may induce to erroneous=20
identifications if people do not expect to find a "=D2deltaic""s=D3 species=
 out=20
of a delta.

=09Concerning the second point quoted from Jim Blakes, we havehere,
in the North Western Mediterranean, a clear example of a species=20
which main mean of transport (and, thus, invasion) are the small boats=20
(particularly, their anchors). This is not a worm nor an animal. Is the=20
tropical alga Caluerpa taxifolia. The origin of the presence of this=20
algae, toxic for most Mediterranean herbivores, has been attributed to an=
=20
escape from an aquarium near the French coasts. Now, the species is=20
quickly increasing its area of distribution. Some Italian and Spanish=20
coasts are affected, together with French coasts, and its expansion can=20
not be controlled by herbivores. In fact, there are no effective grazers=20
of this algae here. It grows quickly, and grows over the typical=20
Mediterranean communities. Particularly negative are their effects on the=
=20
Posidonia oceanica meadows. This endemic species, probably supporting one=
=20
of the most diverse Mediterranean communities, is now being substituted=20
by virtually monospecific beds at the areas =D2colonised=D3 by C. taxifolia=
=20
(no plants nor animals coexist with this species). As C. taxifolia mainly=
=20
occurs in shallow waters, including small sportive harbours, and no=20
sexual reproduction has been reported till now, small boats carrying=20
fragments are the main mechanism of dispersion. And it appears to be very=
=20
effective.

=09Although this last comment does not concern directly to annelids,=20
worms are virtually absent of the bottoms covered by Caulerpa taxifolia=20
beds. Thus, most of the "=D2future generations of north western=20
Mediterranean polychaetologits"=D3 would, why not, becom e =D2future=20
generations of north western Mediterranean specialist on C. taxifolia"=D3.



Daniel Martin

Centre d'=D5Estudis Avacnats de Blanes (C.S.I.C.)
Cami de Santa Barbara s/n, 17300-Blanes (Girona), Catalonia (Spain)

Dani@ceab.es



From BIOSCI-REQUEST  Tue Dec  5 14:28:42 1995
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From: Geoffrey Read <Geoffrey.Read@actrix.gen.nz>
Message-Id: <199512052227.LAA13431@atlantis.actrix.gen.nz>
Subject: ANNELIDA Admin - misdirected unsubscribes
To: annelida@net.bio.net (ANNELIDA list)
Date: Wed, 6 Dec 1995 11:27:03 +1300 (NZDT)
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Folks,

I'd prefer to talk about annelids but:-

There seems to have been a bit of follow-the-leader in the 'unsubscribes' sent
to the ANNELIDA list members rather than the server address. This is of course
ineffectual and a solecism in the eyes of other subscribers. Hence I'm posting
a reminder of how to do it. It should hardly be necessary in a list that
has only been going less than a month. Please DO read and save the
instructions for any list you subscribe to. 

Many people ignore messages without subject headers, unfortunately some
of us don't have that luxury.

If you wish to unsubscribe send the following line in the body of an e-mail to
the *server* address <biosci-server@net.bio.net> 

unsubscribe annelida

For the rest of us -- Send your replies and contributions to the *list*
address <annelida@net.bio.net>. Please check the headers before you send
to be sure your message is going where you want it to. The default for
ANNELIDA is for replies to go to the originator of the message. You must
change the 'to:' line if you intend to make your message public.

By the way the distribution of public messages is exceedingly slow at the
moment. I read Jim Blake's message on the gopher archive hours before I
received it in the mail. I'll check with Bionet whether this can be improved,
or is a "feature" of their server. Meantime a quick look at the gopher will
tell you whether your message is being processed and what else is on its way.

Thanks for listening thus far,


Geoff  
-- 
   Geoff "discussion leader" Read <gread@actrix.gen.nz>
   Annelida resources =>  http://www.actrix.gen.nz/users/chaeto/index.html

From BIOSCI-REQUEST  Wed Dec  6 00:24:17 1995
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From: gbellan@com.univ-mrs.fr
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Date: Wed, 6 Dec 1995 09:27:32 +0100
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Subject: Dani Martinez message

Dear Annelidians

Don't desesperate
Dani Martinez send us a message (more or less a reply to Jim Blake -Salut Jim!)
I read in this message:

This endemic species (Posidonia oceanica), probably supporting one=
>=20
>of the most diverse Mediterranean communities, is now being substituted=20
>by virtually monospecific beds at the areas =D2colonised=D3 by C. taxifolia=
>=20
>(no plants nor animals coexist with this species). 

In fact, I can said there is lot of Polychaetes species (and other
Invertebrates and fishes, and Algae) among Caulerpa taxifolia, even if
diversity and abundance have generally decreased
More details? send an e-mail to  gbellan@com.univ-mrs.fr or
bellan@com.univ-mrs.fr

a small sample of litterature (in French or English):

1993) BELLAN-SANTINI, D., ARNAUD, P., BELLAN, G. & VERLAQUE M.- Résultats
préliminaires sur la faune d'invertébrés du peuplement à Caulerpa taxifolia
des côtes de Provence (Méditerranée nord-occidentale). International
Workshop on Caulerpa taxifolia, GIS Posidonie publ., France .
1994 D. BELLAN-SANTINI, P.M. ARNAUD & G. BELLAN. Affinités entre
peuplements benthiques méditerranéens avec et sans Caulerpa taxifolia. 2nd.
International Workshop on Caulerpa taxifolia, Barcelone, 15-18 décembre
1994, GIS Posidonie publ., France  (under press)
1994) D. BELLAN-SANTINI, ARNAUD, P., BELLAN, G. & VERLAQUE M.- The
influence of introduced alga Caulerpa taxifolia, on the specific diversity
of the marine biota. Marine biodiversity: causes and consequences, York, 30
august- 2 september 1994. 
1995 BELLAN-SANTINI, D., ARNAUD, P., BELLAN, G. & VERLAQUE M. - The
influence of the introduced alga Caulerpa taxifolia on the marine biota. 
J. Mar. Biol. Ass. , .Plymouth, (under press)

Much more has been published by many Mediterranean colleagues (mostly
italian, spanish and french) about Invertabrates, fishes and algae 
Dr. G. BELLAN    
Centre d'Oceanologie de Marseille (U.A. CNRS 41)
Station marine d'Endoume
Rue Batterie des Lions  .
13007 Marseille 
Tel. 91 04 16 12        .
Fax. 91 04 16 35        .
E-mail. gbellan@com.univ-mrs.fr  .






From BIOSCI-REQUEST  Wed Dec  6 04:51:24 1995
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	from CHEMPATH; Wed Dec  6 14:49:16 1995
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unsubscribe annelida


          Eric H.Harley Ph.D.,M.D.
          Dept. of Chemical Pathology
          University of Cape Town         Tel :(021) 406 6222
          Observatory, 7925               Fax :(021) 406 6153
          South Africa             E-Mail : HARLEY@CHEMPATH.UCT.AC.ZA


From BIOSCI-REQUEST  Wed Dec  6 14:35:12 1995
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To: annelida@NET.BIO.NET
From: ROLFP@CLEMSON.EDU
Subject: PREDATORS OF EARTHWORMS

I am searching for a list of all Families of macroinvertebrates and or
amphibians  that feed on earthworms. Does anyone know of such a list?
Also, I have found this this annelida email list useful and plan to
continue as a subscriber, but wonder whether there exists a list for
people of the platyhelminthes persuasion.

From BIOSCI-REQUEST  Wed Dec  6 19:41:59 1995
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Date: Wed, 6 Dec 1995 22:42:24 +0000
To: annelida@net.bio.net
From: fdutra@Nantucket.Net (Frank Dutra)
Subject: Polydora problems

Hello all,

Any information on Polydora sps. (websteri- <1mm?) would be greatly
appreciated. Cultivated scallops (argopecten irradians) have been infected
resulting in some minor mortality and depressed growth.The larger problem
is a lowering of market appeal for infected animals due to unsightly
conchillion deposits. Infection appears to be of a temporal nature,
presumably during the larval stage. Scallops deployed to the field during
August are free of infection, while those deployed earlier are roughly 50
%.I would like to avoid or minimize this problem next year if possible, and
would gladly collaborate (implementation and data collection) with any
experimental procedures suggested.

Another type of infection being reported by processors is a spore like body
(1mm) found in the adductor muscle of commercially harvested scallops.
Infected animals appear in distinct locals independent of substrate or flow
regimes. I vaguely remember some work being done at U-Conn a few years back
as to the identity, vectors, and intermediate hosts of this organism.

Any info or thoughts on the above will be greatly appreciated,..living on
an island has it's drawbacks when trying to access the literature. Thanks,
and keep up the good work!



Frank Dutra
Nantucket Harborlife    (formerly Nantucket Research and Education Foundation)
0 Easton Street
Nantucket, MA  02554

Lab:    (508) 325-7075
Office: (508) 228-5569
FAX     (508) 228-2649                  fdutra@nantucket.net

Frank Dutra
Nantucket Harborlife    (formerly Nantucket Research and Education Foundation)
0 Easton Street
Nantucket, MA  02554

Lab:    (508) 325-7075
Office: (508) 228-5569
FAX     (508) 228-2649                  fdutra@nantucket.net



From BIOSCI-REQUEST  Thu Dec  7 00:15:34 1995
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From: "Helmut Zibrowius" <hzibrowi@com.univ-mrs.fr>
Date: Thu, 7 Dec 95 09:10:25 MST
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To: annelida@net.bio.net
Subject: Martinez' endemic

On Tue, 5 Dec 1995 10:59:03 +0000, Dani Martinez wrote:

>I had the opportunity to describe a
>new capitellid genus from a brackish, shallow-water, enclosed Bay of the
>Ebro's Delta in the Iberian peninsula which I named "Pseudomastus
>deltaicus". It was very abundant there (about 2000 individuals per square
>meter).

REPLY: new species continue to be currently described from the 
Mediterranean. Take the example of the 12th brachiopod species - thus even 
in a group that generally catches attention and that is not speciose in the 
Mediterranean:
  Logan A., Zibrowius H., 1994. A new genus and species of rhynchonellid   
  (Brachiopoda, Recent) from submarine caves in the Mediterranean Sea.  
  Marine ecology [P.S.Z.N.I], 15 (1): 77-88.

Or take the case of a new benthic ctenophore (also a group less speciose 
than Polychaeta):
  Carre C., Carre D., 1993. Ctenella aurantia, genre et espece nouveaux 
  de ctenophore tentacul‚ (Ctenellidae fam. nov.) mediterraneen, sans  
  colloblastes et avec ventouses labiales. Can. J. Zool. 71 (9): 1804-1810.

>As Mediterranean waters have been (and are being) largely explored,
>I thought the species was endemic.

REPLY: That's a rather naive view. For some groups the Mediterranean 
may indeed be better explored than other areas, but that does not garantee
that any new species therein is an endemic. See the above cited cases: the 
authors were cautious enough to not appoint those species endemics.
Given Martinez' strong believe that (nearly) everything is well known in 
the Mediterranean, it would even be more logical to see in each newly 
discovered form a recent immigrant from elsewhere...
There are indeed many aliens of various origins successful in the 
Mediterranean. See the following compilation:
  Zibrowius H., 1992. Ongoing modification of the Mediterranean fauna and 
  flora by the establishment of exotic species. Mesogee [Bulletin du Museum 
  d'histoire naturelle de Marseille], 51, 1991: 83-107.   

>do not know whether this could be a case of recent migration,
>previous misleading identifications, hazard of looking now at the right
>zones, or whatever.

REPLY: Recent migration is another explanation often proposed without 
detailed arguments. But the personal hazard of looking at special biota 
and areas and the ability of being interested in some type of organism may 
generally be a more likely explanation of the discovery.   

>The main point is that the name of the species, directly linked to the type
>locality, may induce to erroneous identifications if people do not expect
>to find a "deltaic" species out of a delta.
REPLY: No, that's really NOT the main point. A name once given with 
reference to the first collecting site can evidently not cover all 
situations to be discovered lateron. All somewhat experienced taxonomists, 
biogeographers, etc. should be aware of that. Nevertheless, names 
totally incorrect with reference to the origin should preferably have been 
avoided: such as Posidonia oceanica (that seagrass appears to be a true 
endemic of the Mediterranean - never duely reported from the - Atlantic - 
Ocean).

  ----------------------------------- 
  Helmut ZIBROWIUS
  (Centre d'Oceanologie de Marseille)
  Station Marine d'Endoume
  Rue Batterie des Lions
  13007 Marseille / France
  TEL: (intern. - 33) 9 1 0 4 1 6 2 4
  FAX: (intern. - 33) 9 1 0 4 1 6 3 5
  E-MAIL: <hzibrowi@com.univ-mrs.fr>
  -----------------------------------

From BIOSCI-REQUEST  Thu Dec  7 08:32:59 1995
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Date: Thu, 7 Dec 1995 17:33:53 +0000
From: Dani Martinez <dani@azathoth.ceab.es>
Subject: Mediterranean diversity
To: annelida@net.bio.net
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Dear colleagues,

I am happy with the responses to my message. It was not intending to be 
catastrophic. I was only a little bit worried about the richness of the 
Mediterranean sea. Now, it seems that we are not in front of a drastic 
reduction, but we are the observers (and, some, also describers) of a change.

Second point. Concerning my coments on the new genus I described. I 
realised some of my points of view were a litle bit naive. I was only trying 
to provide some reflexions to the discussion generated around type 
localities.

Third point. I do not now why, by there is a litle confussion concernig 
my name. It is not "Dani Martinez", but Dani Martin. 

Thank you very much.


-------------------------------------------------------------------
Daniel Martin.
C.E.A.B. (C.S.I.C)
Cami de Santa Barbara s/n, 17300-Blanes (Girona), Catalonia (Spain)
Dani@ceab.es
-------------------------------------------------------------------

From BIOSCI-REQUEST  Fri Dec  8 01:48:11 1995
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From: Geoffrey Read <Geoffrey.Read@actrix.gen.nz>
Message-Id: <199512080946.WAA21890@atlantis.actrix.gen.nz>
Subject: Re: Re, type localities and distributions of species
To: annelida@net.bio.net (ANNELIDA list)
Date: Fri, 8 Dec 1995 22:46:29 +1300 (NZDT)
In-Reply-To: <199512050508.VAA22189@ix8.ix.netcom.com> from "JAMES BLAKE" at Dec 4, 95 09:08:23 pm
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James Blake once wrote:
>So, I ask the question as to what importance is the type locality in 
>understanding a species, if it has been moved around from place to 
>place and the actual area of the world where it originated is somewhere 
>quite different?

Maybe Jim was trolling for bites but here goes ...
 
Type locality has no importance in understanding the biogeography of a 
taxon as long as it is definitely a single species. But if I see a new 
record of 'cosmopolitan' _Paraprionospio pinnata_ outside type locality 
Chile I'm going to take some convincing it's genuine. Also, to state the 
obvious, in the absence of the original specimens, someone goes back to the 
type locality (and hopes they don't find more than one member of the genus, 
and that the site is not now 'infilled' dry land).
 
>I recall a conversation I once had with a colleague who refused to 
>accept an identification I had made because the type locality was 
>somewhere far distant from where the specimens being identified were 
>collected. ...
 
When an expert has done the id there is usually no problem but most 
polychaete ids aren't made by experts. In addition to the basic
nomenclatural role, type localities should have cautionary influence when 
making routine identifications because the overwhelming majority of 
polychaete species have limited distributions. For instance in New Zealand 
we don't especially need any *more* erroneous records of vaguely similar 
species from far-flung countries. 
 
>I think you see the point, namely that type localities are only the
>places where the animals were first collected, not necessarily where
>they are most common or where they originated.
 
Yes. Or hopefully first collected. Sometimes they have been named elsewhere 
too, especially the wandering ones ;-)
 
Geoff
-- 
   Geoff Read <gread@actrix.gen.nz>
   Annelida resources =>  http://www.actrix.gen.nz/users/chaeto/index.html

From BIOSCI-REQUEST  Fri Dec  8 08:17:36 1995
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From: jablake@ix.netcom.com (JAMES BLAKE )
Subject: RE, Paraprionospio pinnata
To: annelida@net.bio.net

Dear annelid workers, 

Geof Read just wrote, 

>Type locality has no importance in understanding the biogeography of a 
>taxon as long as it is definitely a single species. But if I see a new 
>record of 'cosmopolitan' _Paraprionospio pinnata_ outside type 
>locality Chile I'm going to take some convincing it's genuine. Also, 
>to state the obvious, in the absence of the original specimens, 
>someone goes back to the type locality (and hopes they don't find more 
>than one member of the genus, and that the site is not now 'infilled' 
>dry land).

I did a fairly complete review of North American Paraprionospio as part 
of the Taxonomic Atlas of the Santa Maria Basin ..., Vol. 6, the 
spionid chapter.  My conclusion is that the species is present in both 
North and South America, but most certainly does not occur in the 
western Pacific.  I also think it is probably the only species in the 
Americas.  I have not seen specimens from the eastern Atlantic, so 
cannot comment on any records from there. P. pinnata has several unique 
features that make its identification is relatively easy. This will be 
reviewed in my chapter.  Vol. 6 is scheduled for publication in 
February.  More later on vol. 5, out this month.

Jim Blake
Woods Hole, MA
(jablake@ix.netcom.com)

 

From BIOSCI-REQUEST  Fri Dec  8 14:40:02 1995
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From: jablake@ix.netcom.com (JAMES BLAKE )
Subject: Re, Taxonomic Atlas Vol. 5
To: annelida@net.bio.net

Dear Annelid workers, 

On behald of the editors and contributing authors, I pleased to 
announce the publication of Volume 5 of the "Taxonomic Atlas of the 
Benthic Fauna of the Santa Maria Basin and Western Santa Barbara 
Channel."  Of this 14-volume series, the annelids comprise volumes 5-7. 
 

*********************************************

Blake, J.A., B.  Hilbig and P.H. Scott (editors).  1995. Taxonomic 
Atlas of the Benthic Fauna of the Santa Maria Basin and Western Santa 
Barbara Channel.  Volume 5--The Annelida Part 2. Polychaeta: 
Phyllodocida (Syllidae and Scale-Bearing Families), Amphinomida, and 
Eunicida.  Santa Barbara Museum of Natural History.viii + 378 pp. + 
Appendices A-B.  Color cover illustration.  (Date of Publication: 5 
December 1995).

Contents: 

Chapter 1.  Family Syllidae Grube, 1850 (by Jerry D.  Kudenov and  
Leslie H.  Harris): 1-97.

Chapter 2.  Family Aphroditidae Malmgren, 1867 (by James A.  Blake):  
99-104.

Chapter 3.  Family Polynoidae Malmgren, 1867 (by R.  Eugene Ruff): 
105-166.

Chapter 4.  Family Acoetidae Kinberg, 1865 (by James A. Blake): 
167-174.

Chapter 5.  Family Pholoidae Kinberg, 1858 (by James A.  Blake):        
175-188).

Chapter 6.  Family Sigalionidae Kingberg, 1856 (by James A.  Blake): 
189-206.

Chapter 7.  Family Amhinomidae Lamarck, 1818 (by Jerry D.  Kudenov): 
207-215.

Chapter 8.  Family Euphrosinidae Williams, 1851 (by Jerry D.  Kudenov): 
217-228.

Chapter 9.  Family Onuphidae Kinberg, 1865 (by Brigitte Hilbig): 
229-262.

Chapter 10.  Family Eunicidae Savigny, 1818 (by Brigitte Hilbig): 
263-278.

Chapter 11.  Family Lumbrineridae Malmgren, 1867, emended Orensanz, 
1990 (by Brigitte Hilbig):  279-313.

Chapter 12.  Family Oenonidae Kinberg, 1865, emended Orensanz, 1990 (by 
Brigitte Hilbig): 315-339.

Chapter 13.  Family Dorvilleidae Chamberlin, 1919 (by Brigitte Hilbig): 
341-364.

Appendix A.  List of Abbreviations on Figures: 365-366.

Appendix B.  Lists and Maps of Stations:367-372.

Index: 373-378.

This volume represents the second of four planned volumes on the marine 
annelids of the Santa Maria Basin and Western Santa Barbara Channel.  
Volume 5 concludes treatment of the so-called "errant" polychaete 
families.   

A total of 115 species in 57 genera and 13 families are described.  
Chapters are included on the Syllidae, five scale-bearing families, 
Amphinomidae, Euphrosinidae, and five euniciform families.  Eighteen 
species are new to science as follows: Syllidae (12), Polynoidae (1),
Pholoidae (1), Onuphidae (1), Lumbrineridae (1) Oenonidae (1), and 
Dorvilleidae (1). The chapter on the Syllidae represents a major 
revision of some eastern Pacific genera and species of the subfamilies 
Exogoninae, Eusyllinae, and Syllinae.  All 34 species treated are 
described and illustrated.  The 22 species in the chapter on Polynoidae 
include several poorly known species, illustrated in detail for the 
first time.  One new genus and species is described.   For the
Aphroditidae, only juveniles were available in the MMS collections 
precluding any extensive review of the California fauna.  Sigalionidae 
and Acoetidae include previously known species, with updated 
descriptions and keys.  The common amphinomid, Chloeia pinnata was 
found to have unusual bifurcate notosetae in posterior segments.   In 
the Euphrosinidae, two species were identified and described.  In the 
Oenonidae, Arabella pectinata and A. protomutans are newly reported for 
California.  Keys are provided to all species treated in this volume.

This volume may be ordered from the Santa Barbara Musem of Natural 
History, 2559 Puesta del Sol Road, Santa Barbara, California USA 
93105-2936.  Price: $34.00 plus $4.00 shipping, U.S. or $7.00 overseas. 

All payment must be in U.S. dollars.

Contact Dr. Paul H.  Scott (SBMNH) for information about this and other 
volumes in the series (inverts@sbmnh.rain.org). 

+++++++++++++++++++++++++++++++++++++++++++++++

James A. Blake
ENSR, 89 Water Street
Woods Hole, Massachusetts 02543
(jablake@ix.netcom.com)




From BIOSCI-REQUEST  Sun Dec 10 16:19:21 1995
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From: "Petersen, Mary E.       {ZMUC}" <mepetersen@zmuc.ku.dk>
To: ANNELIDA <annelida@net.bio.net>
Subject: RE: Mediterranean diversity & parasites
Date: Mon, 11 Dec 95 01:15:00 DST
Message-ID: <30CB8686@AKI.KU.DK>
Encoding: 73 TEXT
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Hello, Annelidaners!

There is no doubt that even though the Mediterranean is well investigated, 
its biodiversity is greater than presently indicated, with or without 
introduced _Caulerpa taxifolia_.  We can therefore expect a large number of 
new and newly reported taxa as the different groups are critically reviewed 
and revised.

One of the reasons for this is that modern criteria often reveal that what 
earlier was considered INTRAspecific variation is actually INTERspecific 
such.  Moreover, many older species descriptions in reality only describe 
the genera, without sufficient information to permit accurate species 
recognition.  I have found this often in, e.g., _Pholoe_, cirratulids and 
chaetopterids, many of which are very broadly defined.  Most of the 
Mediterranean species I have seen are either undescribed or newly reported, 
yet many of them are hiding under well known names which may belong to 
species not even present in the area.  Within a restricted area it is 
usually not a problem to recognize the individual species present.  The 
difficulties arise when local species must be differentiated from similar 
ones from distant areas.

This brings in the problem of endemic and introduced species.  Having seen 
some Mediterranean cirratulid species that were uncomfortably close to ones 
in other parts of the world, e.g., California, and not knowing if they were 
the same or introduced one place or the other, I came to think of a paper 
presented at the 1989 Long Beach polychaete conference by Thomas G. Douglas 
and Ira Jones (1991. Parasites of California spionid polychaetes. -- Bull. 
Mar. Sci. 48 (2): 308-317).

These authors point out that one of the ways one can recognize recently 
introduced species is that they usually LACK their normal assortment of 
endoparasites, thus the spionid _Pseudopolydora paucibranchiata_ had none, 
suggesting a recent introduction to the coast of southern California.  Among 
the endoparasites present, at least for the gregarines the authors suggested 
that there might be some species specificity and that it might be possible 
to identify the polychaete species from its parasites.

This approach to distinguishing between native and introduced species would 
seem to have an advantage over more traditional electrophoretic techniques 
in that it not only would show a difference (with and without endoparasites) 
but also would make clear which population was more likely to be the 
original one.  The difficulty is that it requires specialist knowledge that 
most of us lack and would have to acquire.

Are there other recent studies on the use of endoparasites for 
differentiating native and introduced populations of a species?

How much do such endoparasites (e.g., gregarines, coccidia, microsporidia, 
mesozoans, bacteria) influence differences obtained by electrophoretic 
comparisons of different populations?  I have not used these techniques, so 
my apologies for unenlightened questions, but I could imagine that the 
presence or absence of certain bands might be due to the presence or absence 
of certain parasites and not because of differences in the polychaetes.

Has anyone done anything with this?  Are there standard techniques for 
cleaning very small specimens before use to minimize or eliminate such 
possible errors?

With greetings from Copenhagen,

Mary
 -------
Mary E. Petersen
Zoological Museum, University of Copenhagen
Universitetsparken 15, DK-2100 Copenhagen O, Denmark
Tel  +45-35 32 10 67     Fax +45-35 32 10 10
E-mail: mepetersen@zmuc.ku.dk
 -----------------------------
Problems with Mediterranean (or other) cirratulids, ctenodrilids or 
chaetopterids?  Contact me at above E-mail for more information.   Mary.
>oooooooo)):>
 ----------------------


From BIOSCI-REQUEST  Mon Dec 11 02:13:18 1995
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X-POPmail-Charset: British
Date: Mon, 11 Dec 95 11:06:11 GMT
From: (Gordon Paterson) <gljp@nhm.ac.uk>
Message-Id: <39971.gljp@nhm.ac.uk>
To: annelida@net.bio.net
Subject: Type nomeclature and rules

I have read Jim Blake's correspondance about types and localities with 
interest. It highlights an important change in our perception about 
species distributions. One of the  orginal reason for a type locality was 
that anyone could then go back to that locality and collect samples of the 
species. It fixed the locality in space. However, in marine systems 
species tend to be widespread and variable in occurrence in time and space 
so the type locality has less relevance. So how should we view the idea of 
a type locality, which under the ICZN rules of nomenclature, is 
considered to be a necessary/ highly desirable part of the information 
about a species?
Personally, I view it as a locality where a group of specimens of the 
species were originally collected, to be considered in the context of the 
overall distribution of the species. A further point about type localities 
is that if you nominate syntypes, instead of a holotype, from a across the 
biogeographic range it gets away from the problems of fixating on a type 
locality. 
Finally, the Commission is about to produce a new edition of the 
rules of nomenclature and would like to hear from anyone who has views on 
the current rules. There is a draft available for comment (they would be 
grateful for contributions of 3 pounds sterling or 5 US dollars to cover 
postage costs, etc.). 
The contact is :Dr Philip Tubbs
Executive Secretary
International Commission on Zoological Nomenclature
The Natural History Museum
London SW7 5BD
e-mail: pkt@nhm.ac.uk
The consultation process will last until May 1996 with the new rules taking 
effect in 1997.
Gordon L J Paterson
Dept. of Zoology
tel: +44 (0)171 938 9414; Fax:+44 (0)171 938 9158 
(drop the zero outside the UK)
E-mail: gljp@nhm.ac.uk

From BIOSCI-REQUEST  Mon Dec 11 05:37:54 1995
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To: annelida@net.bio.net
From: apdl@ozemail.com.au (Andrew Lord)
Subject: Marine worms feed

Dear annelid-o-philes,

I have recently joined Annelida and received information with my
subscription that the main raison d'etre for the group is discussion of
"research on the culture of annelids, but operation of commercial and home
worm-farms will not be a discussion topic".

I am interested in the culture of marine finfish and in the possibility of
using cultured marine worms as a feed during the very early (late larval)
stages of fish growth.

Specifically, I am looking to find if anybody has:

1) cultured marine worms, and if so can they direct me to practical help
with this (people / publications)?
2) used cultured worms as an early feed for finfish?
3) any data on the total fat content and percentage content constituted by
polyunsaturated fatty acids (etc), in marine worms?
4) information on any newsgroup that addresses questions on the commercial
culture of marine worms?


I would very much appreciate any feedback on these questions,


Thankyou in advance,



Andrew Lord


___________________________________________________________
Dr Andrew P.D. Lord
PO Box 142
Belair, SA,
5052
AUSTRALIA
e-mail: apdl@ozemail.com.au
___________________________________________________________



From BIOSCI-REQUEST  Mon Dec 11 05:58:23 1995
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Date: Mon, 11 Dec 1995 08:56:46 -0400
To: annelida@net.bio.net
From: norenbur@onyx.si.edu (Jon Norenburg)
Subject: Re: Type nomeclature and rules

>Finally, the Commission is about to produce a new edition of the
>rules of nomenclature and would like to hear from anyone who has views on
>the current rules. There is a draft available for comment (they would be
>grateful for contributions of 3 pounds sterling or 5 US dollars to cover
>postage costs, etc.).
>The contact is :Dr Philip Tubbs
>Executive Secretary
>International Commission on Zoological Nomenclature
>The Natural History Museum
>London SW7 5BD
>e-mail: pkt@nhm.ac.uk
>The consultation process will last until May 1996 with the new rules taking
>effect in 1997.
In the U.S.A. send $5 to:
Dr. Alan Norrbom, Treasurer
American Association for Zoological Nomenclature
c/o MRC 168
National Museum of Natural History
Smithsonian Institution
Washington, DC 20560
--

Jon L. Norenburg
Department of Invertebrate Zoology-MRC 163
National Museum of Natural History
Washington, DC  20560



From BIOSCI-REQUEST  Mon Dec 11 07:53:40 1995
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From: "Petersen, Mary E.       {ZMUC}" <mepetersen@zmuc.ku.dk>
To: ANNELIDA <annelida@net.bio.net>
Subject: ICZN-4 also available online
Date: Mon, 11 Dec 95 16:50:00 DST
Message-ID: <30CC6171@AKI.KU.DK>
Encoding: 199 TEXT
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Gordon Paterson and Jon Norenberg have just indicated that printed copies of 
the draft of the new International Code of Zoological Nomenclature are 
available for a small fee from the UK and USA.

As many of you already know, this information is also available for free 
online, where ASCII or PostScript copies of the code can be downloaded from 
the following sites as follows:

 -------------- Begin forwarded message 1 --------------------

Return-Path: <owner-ICZN-4@CMSA.BERKELEY.EDU>
Date:         Thu, 15 Jun 1995 08:03:58 -0700
From: "L-Soft list server at UCBCMSA (1.8b)"
    <LISTSERV@CMSA.BERKELEY.EDU>
Subject:      WELCOME to ICZN-4
To: "Mary E. Petersen" <mepetersen@ZMUC.KU.DK>
Reply-To: ICZN-4-Request@CMSA.BERKELEY.EDU
X-LSV-ListID: ICZN-4
 ----------------------------------------------------------------------------  
 --
ICZN-4 International Code of Zoological Nomenclature 4th Edition
                     DRAFT FOR DISCUSSION

WELCOME! This discussion list is dedicated exclusively to the ICZN 4th
Edition Draft for Discussion. The Draft Code and Explanatory Notes can
be obtained via ftp, www, or gopher from the Cornell MUSE Project
Systematics and Biodiversity server (see below for the specific node
addresses of these services).

          WHERE ARE THE DRAFT CODE AND EXPLANATORY NOTES?

The Draft Code itself is available in three formats:

   1) plain ASCII text,
   2) Postscript (very large file!), and
   3) Postscript in a PKZIP file (significantly smaller than the
      raw Postscript version; you need PKUNZIP to decompress it).

MAKE SURE to retrieve the Explanatory Notes also; they contain
important information from the Commission about the Draft Code.
The Notes file is plain ASCII text (called ICZNREAD.ME in the ftp version).

You can obtain the Explanatory Notes and any of the Draft Code formats
using ftp, www, or gopher. Shown below are the Internet node address
(for ftp and gopher), the URL for the www, and the menu choices.

The Postscript format is only useful for printing or viewing on
a system that supports Postscript.

Acknowledgements:
The ICZN Editorial Committee agreed to make these online documents
available for comment.  F. Christian Thompson, USDA, Editorial Committee
member, provided the ASCII text and Postscript versions. Jim Beach,
UCB/MIP, and Julian Humphries, Cornell, made available the MUSE Project
ftp/www/gopher Systematics, Biological Collections and Biodiversity
server, where these files are posted. See below for the BIOSIS server.

Let me know if you find a problem, e.g., corrupted or missing file, with
any of the files. If you have access to ftp, www, or gopher, but need
help to use them, please refer first to your local computer support person.

Peter Rauch peterr@violet.berkeley.edu
 -----------------------------------------------------------------------

Using ftp, go to muse.bio.cornell.edu, login as anonymous,
and change directory (cd) to pub/standards/iczn-4 where the dir command
will show the following files:

 -rw-rw-r--   1 1001     nobody   3439878 Jun  9 15:07 ICZN4PS.ps
 -rw-rw-r--   1 1001     nobody    253171 Jun  9 15:08 ICZN4TXT.txt
 -rw-r--r--   1 1001     nobody    865741 Jun  9 15:09 ICZN4ZIP.zip
 -rw-rw-r--   1 1001     nobody     14982 Jun  9 15:12 ICZNREAD.ME

Before GETting the ICZN4ZIP.zip, make sure to "set binary" transfer mode.
The other files are in plain ASCII "text" mode.

 -----------------------------------------------------------------------

Using a www client, go to URL http://muse.bio.cornell.edu/
and follow the menu choices:

o Standards Organizations: Data Models and Reports
     (DIR) International Code of Zoological Nomenclature

     to arrive at:

          INTERNATIONAL CODE OF ZOOLOGICAL NOMENCLATURE

(FILE) ICZN 4th Edition Explanatory Notes (ASCII text)
(FILE) ICZN 4th Edition Draft for Discussion (PostScript, 3.4 M)
 (BIN) ICZN 4th Edition Draft for Discussion (Postscript, zipped, 800K)
(FILE) ICZN 4th Edition Draft for Discussion (ASCII text, 250K)

 -----------------------------------------------------------------------

Using gopher, go to muse.bio.cornell.edu

and follow the menu choices:

Data Models and Standards Projects/
   International Code of Zoological Nomenclature/

   to arrive at:

           International Code of Zoological Nomenclature

1.  ICZN 4th Edition Explanatory Notes (ASCII text).
2.  ICZN 4th Edition Draft for Discussion (PostScript, 3.4 M).
3.  ICZN 4th Edition Draft for Discussion (Postscript, zipped,.. <PC Bin>
4.  ICZN 4th Edition Draft for Discussion (ASCII text, 250K) .

 -----------------------------------------------------------------------

As a service to the European systematic community, in addition to
retrieving these files from  the muse sites (as detailed above
by Peter Rauch), they can also be accessed via the Zoological
Record gopher and www servers in the UK, as follows:-

Using a www client, go to URL http://www.york.biosis.org/
and follow the menu choice:
    International Code of Zoological Nomenclature


Using gopher, go to  macdui.york.biosis.org
and follow the menu choice:
    International Code of Zoological Nomenclature 4th Edition
        discussion draft

to arrive at:

 1.  ICZN 4th Edition Covering Statement (ACSII text)
 2.  ICZN 4th Edition Explanatory Notes (ASCII text)
 3.  ICZN 4th Edition Draft for Discussion (PostScript, 3.4 M)
 4.  ICZN 4th Edition Draft for Discussion (Postscript, zipped, 800K)
 5.  ICZN 4th Edition Draft for Discussion (ASCII text, 250K)

It should be noted that 1. and 2. are ascii text files as produced by
the Secretariat of the International Commission on Zoological
Nomenclature, London, whereas 3. 4. and 5. are versions prepared in
the US for members and supporters of the American Association for
Zoological Nomenclature to ensure wide and quick dissemination on the
internet, and which may differ in some small respects from the hard
copy version of the draft Code available from the ICZN in London
(contact: The International Commission on Zoological Nomenclature,
c/o The Natural History Museum, Cromwell Road, London SW7 5BD, UK)

Judith Howcroft                          jhowcroft@york.biosis.org
Special Projects Manager
Biosis UK
Garforth House, 54 Micklegate            Tel: +44 (0)1904 642816
York, England, YO1 1LF                   Fax: +44 (0)1904 612793
 ----------------------------------------------------------------------

 ------------- End of forwarded message 1 -----------------

In addition to the above, a special listserve, ICZN-4, has been set up for 
discussion of any and all points of the draft edition.  One may subscribe to 
the listserve as follows:

 ------------- Begin forwarded message 2 --------------------------

Return-Path: <owner-taxacom@CMSA.BERKELEY.EDU>
Message-Id:  <199506142325.QAA19647@violet.berkeley.edu>
Date:         Wed, 14 Jun 1995 16:25:04 -0700
Reply-To: Peter Rauch <peterr@VIOLET.BERKELEY.EDU>
Sender: Biological Systematics Discussion List <TAXACOM@CMSA.BERKELEY.EDU>
From: Peter Rauch <peterr@VIOLET.BERKELEY.EDU>
Subject:      ICZN-4 Discussion List
To: Multiple recipients of list TAXACOM <TAXACOM@CMSA.BERKELEY.EDU>
 ----------------------------------------------------------------------------  
 --
          The ICZN-4 Discussion LISTSERV is now available.

To subscribe, send email to:            LISTSERV@CMSA.BERKELEY.EDU
and include a one-line message saying:  SUB ICZN-4 YourFirstName 
YourLastName

That's all there is to it; you will receive two messages with
information and instructions by return email on how to use ICZN-4.

Availability of the "International Code of Zoological Nomenclature,
4th Edition DRAFT FOR DISCUSSION" online documents was announced
earlier this week here on Taxacom.

ICZN-4 is dedicated _exclusively_ to discussion of the 4th Edition DRAFT
document.

Peter

 -------------------- End of forwarded message 2 ----------------------------


Mary E. Petersen
Zoological Museum, University of Copenhagen
Universitetsparken 15, DK-2100 Copenhagen O, Denmark
Tel MEP direct: +45-35 32 10 67    Fax. +45-35 32 10 10
E-mail:  mepetersen@zmuc.ku.dk
 ----------------------


From BIOSCI-REQUEST  Mon Dec 11 14:34:25 1995
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Date: Mon, 11 Dec 1995 18:20:53 BST
From: Valerie Degas <V.Degas@uea.ac.uk>
Subject: culture of polychaetes
To: annelida@net.bio.net
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Dear Annelidans,


I am probably going to make a laboratory experiment on the regeneration of Scoloplos armiger but 
i have neither references of the culture of polychaetes in laboratory nor on the experiments of 
regeneration of polychaetes.If you have some , and if you are interested in this experiment you 
can contact me whenever you want...
	thanks..

Degas Valerie
postgraduate student
School of biological sciences 
University of East Anglia
NR47TJ Norwich
e-mail : V.Degas@cpca2.uea.ac.uk



From BIOSCI-REQUEST  Tue Dec 12 00:49:22 1995
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From: Geoffrey Read <Geoffrey.Read@actrix.gen.nz>
Message-Id: <199512120847.VAA02630@atlantis.actrix.gen.nz>
Subject: Re: Type nomeclature and rules
To: annelida@net.bio.net (ANNELIDA list)
Date: Tue, 12 Dec 1995 21:47:31 +1300 (NZDT)
In-Reply-To: <39971.gljp@nhm.ac.uk> from "gljp@nhm.ac.uk" at Dec 11, 95 11:06:11 am
Reply-To: gread@actrix.gen.nz
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Gordon Paterson writes:
> ...in marine systems species tend to be widespread and variable in
> occurrence in time and space so the type locality has less relevance.
 
Widespread or ill-diagnosed? For polychaetes the evidence is the latter is 
often the case. In which case type locality might be crucial.
 
> ... A further point about type localities is that if you nominate
> syntypes, instead of a holotype, from a across the biogeographic range it 
> gets away from the problems of fixating on a type locality. 
 
Correct me if I've overlooked something but that appears to be frowned upon in
ICZN-3. "Recommendation 73A. An author who establishes a new nominal
species-group taxon should clearly designate its *holotype*." In addition any
author can subsequently designate one of those biogeographical syntypes as the
lectotype with a type locality.  

There is a lovely subsection on type localities which is apropos on
wandering polychaetes though it might be stretching the intention were it
applied to an established colonising species:

"If captured or collected after being transported by boat, ... , the type 
locality is the place from which the name-bearing type, or its wild 
progenitor, began its unnatural journey."


Geoff
-- 
   Geoff Read <gread@actrix.gen.nz>
   Annelida resources =>  http://www.actrix.gen.nz/users/chaeto/index.html

From BIOSCI-REQUEST  Tue Dec 12 01:04:08 1995
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From: Geoffrey Read <Geoffrey.Read@actrix.gen.nz>
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Subject: Re: Marine worms feed
To: annelida@net.bio.net (ANNELIDA list)
Date: Tue, 12 Dec 1995 22:02:13 +1300 (NZDT)
In-Reply-To: <v01510102acf0a55f9336@[203.7.185.25]> from "Andrew Lord" at Dec 12, 95 00:36:12 am
Reply-To: gread@actrix.gen.nz
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Andrew Lord writes:-
> Specifically, I am looking to find if anybody has:
 
> 1) cultured marine worms, and if so can they direct me to practical help
> with this (people / publications)?
 
Do you have any particular species in mind?
Lots of publications over the years but just recently:
 
Chareonpanich,Charumas; Tsutsumi,Hiroaki; Montani,Shigeru (1994):
Efficiency of the Decomposition of Organic Matter, Loaded on the
Sediment, as a Result of the Biological Activity of Capitella sp. I.
Mar. Pollut. Bull. 28(5), 314-318. [Capitella culture below fish farms]
 
Olive,Peter JW (1994): Polychaeta as a world resource: a review of
patterns of exploitation as sea angling baits, and the potential for
aquaculture based production. Memoires du Museum National d'Histoire
Naturelle 162(Actes de la 4eme Conference internationale des
Polychaetes, Eds: Dauvin,Jean-Claude; Laubier,Lucien; Reish,Donald J),
603-610.
 
Gambi,M Christina; Castelli,Alberto; Giangrande,Adriana; Lanera,P;
Prevedelli,Daniela; Vandini,R Zunarelli (1994): Polychaetes of
commercial and applied interest in Italy: an overview. [ibid.] 593-603.
 
> 4) information on any newsgroup that addresses questions on the commercial
> culture of marine worms?
 
Aquaculture listservs maybe? Try any internet search engine. Also Una 
Smith's 'A Biologist's Guide to Internet Resources' (available at 
sunsite.unc.edu, but not I think updated since last December) mentions 
aqua-l@vm.UOGUELPH.ca, and there are several fisheries lists.
 
> I have recently joined Annelida and received information with my
> subscription that the main raison d'etre for the group is discussion of
> "research on the culture of annelids, but operation of commercial and home
> worm-farms will not be a discussion topic".
 
The limits are flexible but consider mainly the opening sentence of the
charter: "For discussion of the scientific study of Phylum Annelida ...".  We
aim to discuss annelid RESEARCH, into which applied aquaculture & agriculture
with worms might fit, but home gardening, business, and husbandry things that
happen to be associated with worms do not -- in my humble opinion. 

The man who dreamt up the charter, aka ...

Geoff
-- 
   Geoff Read <gread@actrix.gen.nz>
   Annelida resources =>  http://www.actrix.gen.nz/users/chaeto/index.html

From BIOSCI-REQUEST  Tue Dec 12 04:15:19 1995
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From: Benbowme@aol.com
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Message-ID: <951212071313_131163072@emout05.mail.aol.com>
To: V.Degas@uea.ac.uk, annelida@net.bio.net
Subject: Re: culture of polychaetes

We have recently come across some annelids in some freshwater benthic
samples.  These annelids have parapodial-type appendages only on the
posterior portion of the body.  These appendages also seem to lack any
noticeable setae.  We collected these worms along with other polychaetes, but
we have only recently sent them to an expert.  Does anyone have any ideas?
 Any information on identification, references, etc. would greatly be
appreciated.
Thanks
M. Eric Benbow
Dept. of Biology
Univ. of Dayton
513-229-2373  Office
513-229-2021  Fax

From BIOSCI-REQUEST  Tue Dec 12 11:06:13 1995
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From: "Helmut Zibrowius" <hzibrowi@com.univ-mrs.fr>
Date: Tue, 12 Dec 95 13:42:17 MST
Message-Id: <5132.hzibrowi@[139.124.16.1]_POPMail/PC_3.2.2>
Reply-To: <hzibrowi@com.univ-mrs.fr>
X-Popmail-Charset: IBM 8-Bit
To: annelida@net.bio.net
Subject: experts and type localities in polychaetology

In reply to G. Read's discussion contribution of Fri, 8 Dec 1995 22:46:
29 +1300 (NZD), which contained the following partly optimistic remark:

>When an expert has done the id there is usually no problem but most 
>polychaete ids aren't made by experts.... 

   The second part of the sentence is very true and that situation surely 
contributes to increase the general mess.
   Considering the former part, what about those experts like P. Fauvel and 
his priestly school at Angers? They have been and still widely are
considered as experts, especially P. Fauvel, with his very many papers. 
But there are stories about the posterior part of a polychaete described 
as the anterior part of a new taxon and I am inclined to believe those. I 
indeed would have similar anecdotical aspects to add....
   My personal experience with serpulids shows that their identifications 
should always be doubted and verified. Harry ten Hove will probably not 
contradict me?
   We always should be cautious when referring to "expert authorities" 
and always be prepared to play detectives, as in the case of type 
localities: poorly transcribed data, left overs in the dredge from a 
previous haul at a totally different depth, etc. are not uncommon.
   By the way, can somebody tell me where to find Narcon Island?
   That's the type locality of the Antarctic/subantarctic Serpula 
narconensis Baird, from J. Ross' Antarctic exploring expedition. It has 
been quoted many times since 1865.
   Well, I think that I have good reasons to believe that Narcon Island is 
just a poor transcription of Marion Island, a southern island that 
effectively was visited by J. Ross' expedition.
   Play detectives and have fun!

  ----------------------------------- 
  Helmut ZIBROWIUS
  (Centre d'Oceanologie de Marseille)
  Station Marine d'Endoume
  Rue Batterie des Lions
  13007 Marseille / France
  TEL: (intern. - 33) 9 1 0 4 1 6 2 4
  FAX: (intern. - 33) 9 1 0 4 1 6 3 5
  E-MAIL: <hzibrowi@com.univ-mrs.fr>
  -----------------------------------

From BIOSCI-REQUEST  Tue Dec 12 18:07:32 1995
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Received: (from dwkirtley) by igc2.igc.apc.org (8.6.11/Revision: 1.16 ) id SAA10861 for ANNELIDA@net.bio.net; Tue, 12 Dec 1995 18:04:46 -0800
Date: Tue, 12 Dec 1995 18:04:46 -0800
From: "David W Kirtley Ph.D." <dwkirtley@igc.apc.org>
Message-Id: <199512130204.SAA10861@igc2.igc.apc.org>
To: ANNELIDA@net.bio.net
Subject: positions Open: Detective

   ----
Subject: Introduced Sabellariidae
12 December 1995

Artificial introduction of sabellariids

The discussions of artificial ("accidental") introduction of polychaete
species over the past few weeks re-awakened some dim, fuzzy,  thoughts 
I've had about a few of the disjunct, and really enigmatic, patterns of 
distribution of certain sabellariid species that remain after much of 
the "circumtropical," "bipolar," and "comsmopolitan" nonsense is 
dealt with and heavily discounted.

Two examples have clear potential for providing some insight into
"artificially"  introduced populations of sabellariids:

(1)  _Sabellaria vulgaris_ Verrill, 1873, was first described from 
off Massachusetts, where they are extremely abundant.  Morphologic
-ally similar individuals are found in great numbers in nearshore 
areas from Nova Scotia to Cape Kennedy, Florida. They are abundant
in Chesapeake Bay where the find the shells of _Crassostrea_ sp. 
a good place to settle-down and raise a family (Curtis, 1973). 

At a number of localities southward from Cape Hatteras, N. C., _S.
vulgaris_ occurs in clusters of sand tubes with what appear to be 
_S. floridensis_ Hartman, 1944, first described from Englewood, 
on the Gulf coast of Florida.  

They occur in reduced numbers in colonies with _Phragmatopoma 
caudata_ Kroeyer from Cape Kennedy to at least as far south on the 
Atlantic coast as Walton Rocks, just south of Ft. Pierce Inlet 
(Eckelbarger, 1975; 1976)).  

Individuals with virtually the same external appearance are found 
along the northern and western margins of the Gulf of Mexico 
(Kirtley, 1994).  Rioja (1946:196) described some examples from 
Tecolutla, south of Cabo Rojo, Mexico, that may be closely
related forms.

I haven't followed the idea very far but I understand that there
have been a number of cases of transplantation of oysters from 
Chesapeake Bay, and perhaps other localities on the Atlantic Coast
to Louisiana and other Gulf of Mexico sites.  Some of the _S. 
gracilis_ Hartman, 1944, Oakland Airport, San Francisco Bay, 
California) have opercular paleae that are suspiciously similar 
in appearance to those seen in _S. vulgaris_ (See Kirtley,
1994:64:figs. 4.14.1 and 4.14.2). I've heard that "Chesapeake
Bay" oysters have been transplanted to that area in the past.

(2) _Sabellaria nanella_ Chamberlin, 1919, was first described 
from San Francisco Bay.  I am unable to recognize any significant
difference between the holotype and specimens collected from Punta 
Santa Elena, Ecuador; Fortaleza, Ceara, Brazil; and La Pedrera, 
Uruguay.  

Both of these cases may be possibly due to dispersal of sabellariids 
that rode in on the hulls of infested ships, or pirogues, or canoasm
or jangadas.

If anyone would like to share their ideas on this matter, and help
with the detective work (see Zibrowius' comments in this forum, this 
date) I'd be pleased to hear from you...especially if you have some 
Sabellariidae specimens I might see).

Best wishes,

David W. Kirtley, Worm Detective
<dkirtley@igc.apc.org>
Box 2713
Vero Beach, Florida 32961-2713
Phone 407-567-4417
FAX 407-778-2716


From BIOSCI-REQUEST  Tue Dec 12 22:44:22 1995
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From: BIOSCI Administrator <biohelp>
Message-Id: <199512121000.CAA07522@net.bio.net>
To: annelida@net.bio.net
Subject: IMPORTANT: BIOSCI miniFAQ

(LAST REVISION: 08-DEC-95)

This is a new "miniFAQ" designed to answer the questions that come up
the *most frequently*.  The main BIOSCI FAQ (Frequently Asked
Questions) is accessible on the World Wide Web at URL
http://www.bio.net/.

	Contents:
	--------
	1) What to do about "spams," i.e., junk mail, ads, etc.

	2) Examples of subscribing and unsubscribing to the mailing lists.

	3) How to access BIOSCI/bionet newsgroup archives.

	4) The BIOSCI user address and research interest directory.


1) What to do about "spams," i.e., junk mail, ads, etc.
-------------------------------------------------------
BIOSCI is a set of parallel USENET newsgroups (the "bionet" groups)
and mailing lists.  The same postings are distributed on both media
(except for a small number of mailing-list-only groups at
net.bio.net).  Unfortunately it is becoming a despicable practice on
the Internet (by a few people out to make a fast buck) to do automated
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susceptible to this practice, and many spams originate on the USENET
groups and then are passed on to the mailing lists.  However, spammers
also get lists of mailing addresses and hit these too, so neither
medium is immune.

What should you do personally if you get junk mail?
---------------------------------------------------
Just delete it and move on without reading it further.  Filing a
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really understand Internet mail systems, your attempt at protest by
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address of an innocent person that the spammer is victimizing.

What can BIOSCI/bionet do to protect its newsgroups?
----------------------------------------------------
The only solution currently available is to moderate the newsgroup.
If this newsgroup is already moderated, then you are in good shape.
Moderation protects the USENET distribution from about 95% of the
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This takes no more time than that needed to read the message and pass
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Most newsgroups currently have a discussion leader who is responsible
for their newsgroup.  The discussions leaders and their e-mail
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available on the Web at http://www.bio.net/.  If a newsgroup is being
hit with too many junk postings, please contact the discussion leader
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Please do not assume that by simply posting a complaint to the
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to rely on the discussion leaders of each newsgroup to report problems
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We will moderate any of our newsgroups if the discussion leader tells
us that the readership of the group wishes to do so and if a moderator
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entails only a few minutes of work each day.

Moderating a newsgroup will resolve probably 95% of the junk postings
on the USENET distribution.  Unfortunately there are easy ways for
determined spammers to override the moderation mechanism on USENET,
but we can protect our e-mail subscribers from unwanted postings if
the newsgroup is moderated.  We are working on new systems to provide
access to our newsgroups over the WWW.  These will be available by 11
December 1995 and will allow you to use your Web browser to look at
the news postings via our Web site at URL http://www.bio.net.  While
this Web interface will not stop spammers from trying to post to the
groups, this will give you yet another way, besides using USENET news,
to keep the junk out of your personal mail files.  For those of you
with local USENET news systems, the Web interface will also give you
faster access to new newsgroups and recent postings.


2) Examples of subscribing and unsubscribing to the mailing lists.
------------------------------------------------------------------
PLEASE NOTE: The BIOSCI management does NOT act on
subscription/unsubscription requests that are posted improperly to the
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Gory details are in the BIOSCI Information sheets on the Web at
http://www.bio.net.  Below we give an example utilizing the
METHODS-AND-REAGENTS list at both of our two BIOSCI sites:

Users in the Americas and Pacific Rim countries who use the BIOSCI
------------------------------------------------------------------
node at computer net.bio.net:
----------------------------

A) Determine the "listname" which is the <=8 character mail address
                                         ^^^^^^^^^^^^^
   for the group.  These can be found in the BIOSCI Info. Sheet.  For
   the METHODS-AND-REAGENTS group the mailing address is
   methods@net.bio.net.  The listname is the portion of the address to
   the left of the @ sign, i.e., "methods".  The listname is used with
   the "subscribe" and "unsubscribe" commands illustrated below.

B) Mail all commands in the body of a mail message addressed to
   biosci-server@net.bio.net.  Do NOT send commands to the newsgroup
   posting addresses!  Leave the Subject: line blank, any text on it
   will be ignored.

C) In the body of your message put one or more of the following
   commands with an "end" command on the last line, e.g.,

   subscribe methods
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   end

   Do NOT put your e-mail address or other text on these lines.  The
   server only allows you to cancel your subscription if the address
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   Please ask for help at biosci-help@net.bio.net if your address has
   changed, e.g., if you know you are on the list but the server tells
   you that you are not a member.


Users in Europe, Africa, and Central Asia who use the BIOSCI node at
--------------------------------------------------------------------
computer daresbury.ac.uk (also known as dl.ac.uk):
-------------------------------------------------

To subscribe and unsubscribe to/from the BIOSCI lists, you need to
specify the full USENET newsgroup name with "bionet-news." prepended.
The USENET newsgroup names are listed in the BIOSCI Information sheet
on the Web at http://www.bio.net/.  For the METHODS-AND-REAGENTS list
the USENET newsgroup name is bionet.molbio.methds-reagnts, thus the
appropriate commands are

    sub bionet-news.bionet.molbio.methds-reagnts

    unsub bionet-news.bionet.molbio.methds-reagnts

These commands are included in a message addressed to mxt@dl.ac.uk,
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the body of the message as text on the Subject: line is ignored.

To unsubscribe from all the lists at the UK node, use

    unsub bionet-news

Please note that if the address in the list is different than the one
in your mail message header, you will not be able to unsubscribe by
this method. If you have problems, please mail biosci@daresbury.ac.uk.


3) How to access BIOSCI/bionet newsgroup archives.
--------------------------------------------------
Back postings of all BIOSCI/bionet newsgroups can be found on the
World Wide Web at URL http://www.bio.net/.  There are several
searchable newsgroup indices at this site.  E-mail users can search
the BIOSCI archives by using our waismail e-mail server.  For
instructions send the message

help

to waismail@net.bio.net.  Leave the Subject: line blank (anything
entered on the Subject: line is ignored).


4) The BIOSCI user address and research interest directory.
-----------------------------------------------------------
Please take this opportunity to add your name, address, and research
interest information to the BIOSCI User Address Database if you have
not already done so.

You can fill out the address form directly through our Web page at URL
http://www.bio.net/adrform.html.

The address database is reindexed nightly for WWW access (the URL is
http://www.bio.net/).  If you are not directly on the Internet but can
reach it by e-mail, please use our waismail server to access the user
directory.  waismail use is described above.  You can also request a
user address form by e-mail from biosci-help@net.bio.net.

Please check your database entry from time-to-time to see if your
address information is still up-to-date.  Because of our limited
personnel resources, we ask that you resubmit a *complete* form to
revise your entry; we only replace complete entries and do not have
resources to edit old forms.

				Sincerely,

				Dave Kristofferson
				BIOSCI/bionet Manager

				biosci-help@net.bio.net

From BIOSCI-REQUEST  Tue Dec 12 23:08:46 1995
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From: "Helmut Zibrowius" <hzibrowi@com.univ-mrs.fr>
Date: Wed, 13 Dec 95 08:03:54 MST
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Reply-To: <hzibrowi@com.univ-mrs.fr>
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To: annelida@net.bio.net
Subject: travelling polychaetes

In reply to worm co-detective Kirtley's remarks of Tue, 12 Dec 1995 18:
04:46 -0800:

   I cannot help with the sabellariids. I just wish to confirm that 
Crassostrea virginica has been an efficient vector for many species: 
from Atlantic N America to Pacific N America, and also to W Europe. I would 
expect that close to intertidal sabellariids are likely to survive such 
oyster trips. Later, the Japanese oyster Crassostrea gigas carried plenty 
of Far Eastern species to Europe, negligence of the French Fisheries etc. 
authorities helping. Including the serpulid Hydroides ezoensis - also a 
tube worm and thus specially adapted to travelling. Various cases of the 
"Jap-fauna" are mentioned in my compilation on the aliens in the 
Mediterranean Sea. Sabellariids apparently were not among the epifauna of 
batches examined in the 70s, by Y. Gruet. Perhaps they preferred to take a 
later train...

  ----------------------------------- 
  Helmut ZIBROWIUS
  (Centre d'Oceanologie de Marseille)
  Station Marine d'Endoume
  Rue Batterie des Lions
  13007 Marseille / France
  TEL: (intern. - 33) 9 1 0 4 1 6 2 4
  FAX: (intern. - 33) 9 1 0 4 1 6 3 5
  E-MAIL: <hzibrowi@com.univ-mrs.fr>
  -----------------------------------

From BIOSCI-REQUEST  Wed Dec 13 03:17:27 1995
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To: annelida@net.bio.net
From: hove@bio.uva.nl (Harry A. ten Hove)
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Subject: expert taxonomists identifications
Message-ID: <1393265381-2628529@bio.uva.nl>

In reply to G. Read's discussion contribution of Fri, 8 Dec 1995 22:46:
29 +1300 (NZD), which contained the following partly optimistic remark:

>When an expert has done the id there is usually no problem but most 
>polychaete ids aren't made by experts.... 

Challenged by Helmut Zibrowius, I endorse his views on "expert
identifications". I will even go further than he. Any identification is a
product of its epoch. Of course, when it is generally accepted that a
species is occurring world-wide (like "Serpula vermicularis" during the
last few decades), an identification of a large Serpula will end up with
the nominal taxon vermicularis. Nevertheless, from a biogeographic
perspective this is unlikely, and really looking at the material it is
evident that a complex of species is involved.  

Most "experts" have more material than they can handle, and tend to skip
corners whenever possible. A simplified example, which nevertheless may be
valid. The genus Filogranella was described in 1979. Its tube may show 5
longitudinal ridges, not unlike those in the serpulid Vermiliopsis. In
other characters there are some superficial similarities as well.
Preliminary identifications of this group in the field and during visits to
other musea were done upon the supposedly fairly characteristic tube
mainly, so any such identification from before 1979 only could have been
Vermiliopsis, but may include specimens of the untill than undescribed
Filogranella. Between 1979 and 1985 I used two characters for these hasty
field identifications: non-operculate, five keels = Filogranella;
operculate, five keels = Vermiliopsis. Around 1985 I found operculate
representatives of Filogranella, thus operculate specimens of "Vermiliopsis
spec." det. ten Hove 1979-1985 still might include Filogranella. Nowadays I
routinely use the uncini (specialised thoracic chaetae) in addition to
discriminate between the two genera, but I don't have the time to go back
upon the hundreds of previously identified samples. I don't think that many
mistakes like described above actually will have been made, the example
after all was simplified and I often do use not mentioned characters as
well, but cannot exclude the possibility either.

In my 30 years of serpulid experience, I have re-identified material of
almost all broad polychaete specialists like Augener, Day, Fauvel, Hartman,
Hartmann- Schroeder, McIntosh, Monro, Pettibone, Rioja, (and I certainly
are forgetting a few names), as well as that of more specializing, esteemed
colleagues like Bailey-Brock, Ben-Eliahu, Imajima, Knight-Jones, Pillai,
Zibrowius (and if I have omitted someone that does not mean that (s)he is
not esteemed nor that (s)he did not make mistakes), and last but certainly
not least ten Hove. I can assure that none of us is infallible. Some made
more mistakes than others, not every-one of us will have been a phylum
wrong in her/his identification (like Fauvel and Augener), not everyone
will use a book on the Mediterranean fauna for tropical material like
Fauvel did in his Fauna of India, but we all are overburdened with routine
identifications and will slip sometimes.

In conclusion, if "usually" would be defined as "in 80% of the cases", I
might agree with our discussion leader's statement >When an expert has done
the id there is usually no problem< 

I subscribe the intention of his remark
>but most polychaete ids aren't made by experts.... 
but see this as a challenge to us "experts" to produce more and better
review papers, or even better both papers and digitalized data-banks,
giving the non-experts better tools to unburden us of routine ids.
Wormly,
Harry A. ten Hove
Institute for Systematics and Populationbiology
Zoological Museum, University of Amsterdam
POB 94766, 1090 GT AMSTERDAM


tel. 3120 5256906
fax. 3120 5255402
Email: hove@bio.uva.nl


From BIOSCI-REQUEST  Wed Dec 13 08:18:54 1995
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Date: Wed, 13 Dec 1995 08:14:06 -800 (PST)
From: Mike Satterwhite <msatterw@lcsc.edu>
Subject: Re: travelling polychaetes
To: Helmut Zibrowius <hzibrowi@com.univ-mrs.fr>
Cc: annelida@net.bio.net
In-Reply-To: <2301.hzibrowi@[139.124.16.1]_POPMail/PC_3.2.2>
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As a relatively quiet observer of the travelling discussion I'd like to 
point out the obvious. There is some good material being assembled here 
chronicling potentially evolutionarily significant events. It would be a 
good idea if some of you were archiving these communications for future 
reference and perhaps even pattern analysis.

MS
============================================================================
D. Michael Satterwhite, PhD.			Phone: 	208-799-2890 at LCSC
Division of Natural Science and Mathematics	Home:   208-746-3628/7288
Lewis-Clark State College			Fax:    208-799-2064  
500 8th Street
Lewiston, Idaho  83501				e-mail: msatterw@lcsc.edu


From BIOSCI-REQUEST  Wed Dec 13 08:39:05 1995
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From: jablake@ix.netcom.com (JAMES BLAKE )
Subject: Re, Expert Taxonomic Identifications
To: annelida@net.bio.net

Dear Annelid enthusiasts, 

The recent postings by Read, Zibrowius, and Ten Hove on their views 
about "expert" and "non-expert" identifications need to be tempered 
with the point that our knowledge of speciation in polychaetes has 
changed considerably in recent years.  SEMs were not available to most 
of our predecessors; there was little appreciation of how species were 
moved around my man's activities, and there was no knowledge of the 
genetics of sibling species.  We are now examining species at a 
different level of discrimination than we were even 10 years ago.  

Twenty years ago, almost any cirratulid found with cinctures of spines 
encircling its posterior end would probably have been referred to 
Chaetozone setosa with little thought.  

For example, in 1968 I participated in a cruise to the eastern Canadian 
Arctic and in the results (Blake and Dean, 1973: Bull. So. Calif. Acad. 
Sci. 72), did indeed identify such a cirratulid from off Baffin Island 
as Chaetozone setosa. 

In a recent revision of cirratulids from the northeastern Pacific, I 
have identified no less than 16 species of Chaetozone, none of which 
are the real C. setosa.  

A reexamination the above mentioned Arctic "C. setosa" mentioned in the 
1973 paper indicates that it is yet another undescribed species. 

We are going to find this level of speciation in cirratulids all over 
the world and the difficulty of documenting this diversity will be very 
difficult, but it non-the-less exists; and in other families as well. 

My point is that we should not dwell on who did what kind of 
identifications or how well or how poorly.  We all make mistakes, but 
given the level of our knowledge and understanding and what is at our 
disposal at the time, we do the best we can.

Jim Blake
ENSR
89 Water Street
Woods Hole, MA 02543
(jablake@ix.netcom.com)






From BIOSCI-REQUEST  Thu Dec 14 00:29:25 1995
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From: "Helmut Zibrowius" <hzibrowi@com.univ-mrs.fr>
Date: Thu, 14 Dec 95 09:24:31 MST
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To: annelida@net.bio.net
Subject: impact of old unreliable identifications

On Wed, 13 Dec 1995 08:36:58 -0800, JAMES BLAKE wrote:

>The recent postings by Read, Zibrowius, and Ten Hove on their views 
>about "expert" and "non-expert" identifications need to be tempered 
  ......
>My point is that we should not dwell on who did what kind of 
>identifications or how well or how poorly.....

The point simply is that we should be ready to be cautious and critical. 
Those old data representing certain periods (and some people have 
always been more behind the generally admitted period than others) are 
frequently reused in "modern" contexts. Many people now have computers, 
feed in whatever data, and are pleased to have nice graphs drawn, whatever 
insignificant they may be given the input. I do not need to refer specially 
to Polychaeta. For example, I also remember two recent MS on Anthozoa. 
Computer games with whatever data are a general trend.

  ----------------------------------- 
  Helmut ZIBROWIUS
  (Centre d'Oceanologie de Marseille)
  Station Marine d'Endoume
  Rue Batterie des Lions
  13007 Marseille / France
  TEL: (intern. - 33) 9 1 0 4 1 6 2 4
  FAX: (intern. - 33) 9 1 0 4 1 6 3 5
  E-MAIL: <hzibrowi@com.univ-mrs.fr>
  -----------------------------------

From BIOSCI-REQUEST  Thu Dec 14 00:36:35 1995
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From: "Craeymeersch, Johan" <craeymer@cemo.nioo.nl>
To: ANNELIDA <annelida@net.bio.net>
Subject: Marenzelleria viridis
Date: Thu, 14 Dec 95 09:18:00 PST
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The North Amercian spionid _Marenzelleria (Scolecolepides) viridis_ 
(Verrill, 1873) was first reported in European waters in 1982 (McLusky et 
al., 1993). Since then, the polychaete has spread into several estuaries 
around the North Sea and also appeard in the Baltic Sea. Essink and Kleef 
(1993) formulated a hypothesis regarding the possible mode by which the 
spionid polychaete was able to spread.: a second introduction in the Ems 
estuary; and the anti-clockwise water circulation pattern from the two sites 
of introduction (Forth estuary, Ems estuary). Given the genetic differences 
between the North Sea and Baltic populations, the Baltic populations 
probably result from a third introduction (Bastrop et al. 1995).

Last week, specimens of this polychaete were found in the south-west of the 
Netherlands. The benthic fauna of this area, built op by several estuaries, 
was studied intensively by Wolff (1973). Since then the area has been 
changed profoundly my major coastal engineering projects. Several  estuaries 
have been dammed and in the Oosterschelde a storm-surge barrier has been 
built. As a result, some of the estuaries were turned into non-tidal lakes 
or lagoons filled with brackish or saline water.

We might expect Marenzelleria to start colonizing the whole area. In the 
southern Baltic it has become the dominant macrobenthos species (Zettler et 
al., 1995), and given the different water bodies in the Delta area, this may 
happen here as well. Therefore, I am looking for references on the 
distribution of this species in America, and on the ecology of this species.

Thanks in advance,

Johan Craeymeersch
Netherlands Institute of Ecology
Centre for Estuarine and Coastal Ecology
Vierstraat 28
NL-4401 EA Yerseke
the Netherlands

craeymer@nioo.nl






From BIOSCI-REQUEST  Thu Dec 14 10:52:02 1995
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To: annelida@net.bio.net
From: Angel de leon <jadeleon@ccr.dsi.uanl.mx>
Subject: polychaete distribution

Estimados estudiosos en anelidos

   In reply to the discution of the worldwide distribution of some
polychaete species. I found a article titled "Ecological Roulette: The
Global Transport of nonindigenous marine organisms" Science, 261: 78-82
(1993). Cartlon and Geller sampled ballast water from 159 cargo ships in
Coos Bay, Oregon, USA. The ships and their ballast water were originated
from 25 Japanese Ports. They found a minimum of 367 different marine taxa,
including spionids, polynoids among other polychaete families.
   These study may explain the transpacific distribution in the recent past.
The interesting point is, how many transpacific species Jim Blake found in
his Atlas?.

Angel de Leon
Lab. Zoologia de Invertebrados
Fac. Ciencias Biologicas, U.A.N.L.
Ap. Postal 5, Suc. "F"
San Nicolas de los Garza, N.L.
66451 MEXICO

e-mail: jadeleon@ccr.dsi.uanl.mx


From BIOSCI-REQUEST  Thu Dec 14 13:07:28 1995
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From: jablake@ix.netcom.com (JAMES BLAKE )
Subject: Re, Polychaete Distribution
To: annelida@net.bio.net

Dear Annelid enthusiasts, 

Angel de leon wrote:

>   In reply to the discution of the worldwide distribution of some
>polychaete species. I found a article titled "Ecological Roulette: The
>Global Transport of nonindigenous marine organisms" Science, 261: 
>78-82(1993). Cartlon and Geller sampled ballast water from 159 cargo 
>ships in Coos Bay, Oregon, USA. The ships and their ballast water were 
>originated from 25 Japanese Ports. They found a minimum of 367 
>different marine taxa, including spionids, polynoids among other 
>polychaete families.

>   These study may explain the transpacific distribution in the recent 
>past. The interesting point is, how many transpacific species Jim 
>Blake found in his Atlas?.

The cited paper is one of several authored by Jim Carlton detailing his 
interesting work on transport of invertebrates.  An early summary, 
relevent to California appeared in the 1975 edition of Light's Manual. 

The answer to Angel's question, however, is easy if we are considering 
introduced or non-indigenous species.  None!!

The Atlas I am referring to deals almost exclusively with polychaetes 
from the continental shelf and slope, not from estuarine habitats where 
the opportunitistic species found in ballast water colonize.  

Geologically, the estuaries of California are so young, that there are 
relatively few endemic invertebrates.  This provides opportunities for 
invading species that are transported to these estuaries in ballast 
water or through aquaculture transplants.  Typically such species are 
tolerant to shifts in salinity and temperature.  This is not true for 
offshore species found in the deeper waters of the shelf and slope.  
These species have a narrow range of tolerance and occur there because 
of natural transport and colonization.  

Jim Blake
ENSR
89 Water Street
Woods Hole, MA 02543
(jablake@ix.netcom.com)


From BIOSCI-REQUEST  Thu Dec 14 17:19:19 1995
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From: Geoffrey Read <Geoffrey.Read@actrix.gen.nz>
Message-Id: <199512150117.OAA28152@atlantis.actrix.gen.nz>
Subject: Re: Re, Polychaete Distribution
To: annelida@net.bio.net (ANNELIDA list)
Date: Fri, 15 Dec 1995 14:17:13 +1300 (NZDT)
In-Reply-To: <199512142105.NAA15035@ix3.ix.netcom.com> from "JAMES BLAKE" at Dec 14, 95 01:05:14 pm
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JAMES BLAKE writes:-
> Angel de leon wrote:
> >   These study may explain the transpacific distribution in the recent 
> >past. The interesting point is, how many transpacific species Jim 
> >Blake found in his Atlas?.

> The answer to Angel's question, however, is easy if we are considering 
> introduced or non-indigenous species.  None!!

Aglaophamus verrilli (McIntosh, 1885), type locality Marlborough Sounds, 10
faths, New Zealand, is reported from 100-140m depth in the Basin by Brigitte
Hilbig in vol 4. (Other authors have reported it from USA before Brigitte I
should add.) It's quite a distinctive species, but perhaps this is misleading
(like Chaetopterus) and future workers with yet more advanced 'tools' will
manage to separate the Californian version again. However, other NZ marine
fauna have apparently reached West Coast USA across the Pacific (a nudibranch
was it recently?), and, as Jim indicates, the shallow water is the area where
intrusions are most likely. For what it's worth A. verrilli in NZ is a
near-shore species.  

Nephtyid experts please, -- the floor is yours to knock this one on the head
... 

Geoff
-- 
   Geoff Read <gread@actrix.gen.nz>
   Annelida resources =>  http://www.actrix.gen.nz/users/chaeto/index.html

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Date: Thu, 14 Dec 1995 18:51:40 -0800
From: "David W Kirtley Ph.D." <dwkirtley@igc.apc.org>
Message-Id: <199512150251.SAA19017@igc2.igc.apc.org>
To: annelida@net.bio.net
Subject: estuarine evolution


Jim Blake writes:

<<Geologically, the estuaries of California are so young, that there are 
<<relatively few endemic invertebrates.

This statement needs to be clarified a bit, I believe, before it 
becomes generally and uncritically accepted, embedded, and perfused 
through the literature.

Some of the present day estuaries may have been at their present 
positions for relatively brief (geologic) time spans, indeed, but 
there is little reason to imagine that there were no antecedent estuarine
habitats along the California coast that occupied laterally contiguous
positions during lower, or higher, relative sea level stands.  

Whether as the result of global glacio-eustatic rise and fall of sea-
level or local tectonic vertical displacement and/or lateral translation
of fault blocks, the size and shapes and areal extent of the estuarine 
habitats may have changed greatly  from time to time, but this would not 
completely obviate the opportunity for the evolution of endemic 
invertebrates in those estuarine habitats. 

If there is evidence that at any time the California coastal margin was 
a high, sheer, strait cliff, and all the rivers and streams were dammed 
up so that there was no fresh water flowing into the ocean, then my
argument would be baseless and my conclusions dead wrong.

I think I'll e-mail it out and find out, pronto!

Detectively,

David W. Kirtley
<dwkirtley@igc,apc.org>


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Geoffrey Read wrote:

> However, other NZ marine > fauna have apparently reached West Coast USA
across the Pacific (a nudibranch > was it recently?), and, as Jim
indicates, the shallow water is the area where > intrusions are most
likely. For what it's worth A. verrilli in NZ is a > near-shore species. 
> 

With apologies to the group, I will mention a non-annelid here.  A recent
New Zealand arrival to the California coast (although not the one to which
Geoff referred perhaps) is the cephalaspid gastropod Philine auriformis. 
This guy was first reported in Calif by Gosliner (1995) from San Francisco
Bay where it became established within the past decade. SF Bay is, of
course, the typical setting for such invasions, as Jim Blake suggested. 
However it seems not to have been satisfied with confinement to the Bay
and has appeared on the open coast in southern California within the past
year.  It is now being taken routinely in quarterly sampling (both trawls
and benthic grabs) from the Los Angeles region to as far south as San
Diego in depths from 20 to 305 meters.  On the Palos Verdes Shelf near Los
Angeles several hundred have been taken in a ten minute trawl with a small
otter trawl.  Jim Blake contrasted shelf and slope depth communities to
those of estuaries, citing the former's relative immunity to invasion by
non-native species.  Philine auriformis appears to be an exception to the
general case, although its foothold in a hospitable estuary may have
facilitated its subsequent invasion of the shelf and slope.  Another
possibility is an introduction directly to the southern Californian shelf and
slope.

Dave Montagne
Marine Biology Lab
County Sanitation Districts of LA County
mblcsdla@netcom.com


From BIOSCI-REQUEST  Thu Dec 14 20:46:25 1995
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Date: Thu, 14 Dec 1995 20:44:53 -0800
Message-Id: <199512150444.UAA01680@ix2.ix.netcom.com>
From: jablake@ix.netcom.com (JAMES BLAKE )
Subject: Estuarine evolution
To: annelida@net.bio.net

Dear Annelid enthusiasts, 

I am delighted at the various directions discussions are taking with 
regard to transport of exotic species. 

David Kirtley writes in response to my comment about the relatively 
young geological age of California estuaries.

>Some of the present day estuaries may have been at their present 
>positions for relatively brief (geologic) time spans, indeed, but 
>there is little reason to imagine that there were no antecedent 
>estuarine habitats along the California coast that occupied laterally 
>contiguous positions during lower, or higher, relative sea level 
>stands.  

>Whether as the result of global glacio-eustatic rise and fall of sea-
>level or local tectonic vertical displacement and/or lateral 
>translation of fault blocks, the size and shapes and areal extent of 
>the estuarine habitats may have changed greatly  from time to time, 
>but this would not completely obviate the opportunity for the 
>evolution of endemic invertebrates in those estuarine habitats. 

>If there is evidence that at any time the California coastal margin 
>was a high, sheer, strait cliff, and all the rivers and streams were 
>dammed up so that there was no fresh water flowing into the ocean, 
>then my argument would be baseless and my conclusions dead wrong.

If you look at a map of the coastline from the Columbia River to the 
southern California Bight, you will not find very many embayments.  
Most of the coastline is composed of cliffs with debris from former 
cliffs forming extensive rocky areas just offshore.  This has created 
habitat for a wonderful rocky intertidal and subtidal fauna anf flora, 
much of it endemic and well studied.  

Most of the literature dealing with introduced invertebrates in 
California estuaries comes from San Francisco Bay, an estuary that has 
been extensively modified since siltation caused by the hydraulic gold 
mining in the 1850's and thereafter.  Extensive habitat alteration from 
that area and of subsequent land filling operations would most 
certainly have altered endemic faunas.  

However, evidence of a recent geological origin is quite evident in 
Tomales Bay where I worked throughout the 1970's.  This bay is the 
sunken rift zone of the San Andreas Fault and we were always dreading 
the day the "big one" would hit.  According to predictions, the City of 
Los Angeles will one day lie just offshore of my former laboratory, the 
Pacific Marine Station.  Who knows how many "endemic" species and 
polychaete biologists will have come and gone when that time comes. 

>From a transplanted, 3rd generation native Californian, now shoveling 
snow from the first storm of the winter in Massachusetts.

Jim Blake
ENSR
89 Water Street
Woods Hole, MA 02543



From BIOSCI-REQUEST  Thu Dec 14 23:35:47 1995
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From: "Helmut Zibrowius" <hzibrowi@com.univ-mrs.fr>
Date: Fri, 15 Dec 95 08:31:10 MST
Message-Id: <2002.hzibrowi@[139.124.16.1]_POPMail/PC_3.2.2>
Reply-To: <hzibrowi@com.univ-mrs.fr>
X-Popmail-Charset: IBM 8-Bit
To: annelida@net.bio.net
Subject: Re: estuarine evolution

On Thu, 14 Dec 1995 18:51:40 -0800, David W Kirtley Ph.D. wrote:
>.......
>This statement needs to be clarified a bit, I believe, before it 
>becomes /// generally and uncritically accepted, embedded, and perfused 
>through the literature ///.

   Well and pertinently formulated again!
   As for the arguments concerning the non-absence of Californian 
estuaries in former ages, we have a somewhat similar situation in our 
area (NW Mediterranean). Even though the submarine caves we presently 
know (among many others: paleolithic Cosquer cave with entrance at now
-37m; cave with the carnivorous sponge Asbestopluma and hexactinellid 
Oopsacas at ca -20m -- you may have heard about these through TV and 
Nature) had been totally immerged during low stands of the sea level, there 
still remained other (deeper) caves submerged since karstic lime stone 
formations extend into much deeper water, as has been documented by 
submersible dives. At least in that case evidence can be seen even now. 
Submarine caves here are NOT new Holocene biota either.

  ----------------------------------- 
  Helmut ZIBROWIUS
  (Centre d'Oceanologie de Marseille)
  Station Marine d'Endoume
  Rue Batterie des Lions
  13007 Marseille / France
  TEL: (intern. - 33) 9 1 0 4 1 6 2 4
  FAX: (intern. - 33) 9 1 0 4 1 6 3 5
  E-MAIL: <hzibrowi@com.univ-mrs.fr>
  -----------------------------------

From BIOSCI-REQUEST  Fri Dec 15 00:42:05 1995
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