DNA Structure Puzzle #7: Reply to Andrew Dalke

Clive Delmonte clived at ndirect.co.uk
Thu Apr 1 13:07:13 EST 1999


Andrew,    Thankyou for your message dated 31st March.   I don't scan
dejanews so I didn't know there was a part of our dialogue recorded there.
I'm unclear as to your allusion to my not addressing the objections you
posted there.   However, I did respond to your points via the newsgroup on
18th February, and I now enclose a copy of the posting and the attachment.

I have renumbered the Puzzles to give added coherence to the themes, and I
shall be adding new Puzzles to do with Protein-DNA interactions.

As you know, my contention is that one simplifying proposition, a true
side-by-side DNA duplex allows a natural explanation of the experimental
results I cite, all of which are drawn directly from other research reports
and are often supported by quotations taken from those studies.

Apart from Puzzle #7, which attracted your interest last time (as Puzzle
#6), what explanation do you offer for any of the other Puzzles ?


Clive Delmonte
-------------------------------------------------------------
From: "Clive Delmonte" <clived at ndirect.co.uk>
Newsgroups: bionet.biophysics,bionet.xtallography
References: <36b81714.0 at news.netdirect.net.uk>
<Pine.SOL.3.96.990203105728.9386C-100000 at taurus.cus.cam.ac.uk>
<36BE3682.977F323B at bioreason.com> <36beffa3.0 at news.netdirect.net.uk>
<36BF48D4.B38DC74E at bioreason.com> <36c1be39.0 at news.netdirect.net.uk>
<36C249C8.BAF3AED2 at bioreason.com> <36c7fae5.0 at news.netdirect.net.uk>
<36C8F645.3DD3F36C at bioreason.com>
Subject: Re: Is ignorant rambling, was: DNA Structure: An Age of
Refinement...
Date: Thu, 18 Feb 1999 18:24:19 -0000
Lines: 6565
X-Newsreader: Microsoft Outlook Express 4.72.3110.5
X-MimeOLE: Produced By Microsoft MimeOLE V4.72.3110.3

Hello Andrew !

Thankyou for your recent posting.  It is a real pleasure to be able to
discuss this important issue, the nature of the true, normal structure of
duplex DNA, upon which there are genuine differences of opinion.  Indeed, I
have received these comments on the true side-by-side model:

"...I find much of Delmonte's critique of other workers sound enough to
raise doubt in my mind about the bulk of the classical work in this area..."
Prof Steven Benner, ETH Zurich,     and

"...the widely accepted Watson-Crick model is inadequate to explain many
important pieces of data, and, in some cases, defies intuitive biological
and physical logic as a predictive model...I commend you on your recognition
of inconsistencies in the story of DNA..."
Prof Robert Hopkins, University of Houston

Of course, science is not a matter of counting votes, but these comments
illustrate how it is possible for scientists to view the same situations
quite differently.

I'm sorry you were unable to open the WORD attachment containing details of
the model.  With my software, and limited scanning exporting skills, I can
only manage a few formats and the attachment herewith is in *.bmp.  If that
is not OK for you, I would be very happy to send a hard copy to your postal
address, and to those of any other subscribers similarly placed.

Returning to the detail of your last posting, on the thermodynamics side, we
would keep in mind, of course, that a chemical system with a low energy
state may not actually to be able to reach that state under ambient
conditions.  The low energy state may just be inaccessible under the normal
conditions in which the equilibrium is found.   If the double helix were
more stable, as predicted by force field calculations, the double helix,
consisting of thousands of plectonaemic full turns, would not necessarily
exist in vivo.

Your point that force field energies give results reasonably close to those
observed experimentally reminds me that, in using some techniques at least,
such as NMR for distances between resonating nuclei in DNA, it is necessary
to calibrate a number of constants against predictions using the very model
which is being explored.  My knowledge of the process of molecular dynamics
is too poor to know whether there might be any circularity in the
calculations on DNA double helix energies which would, if there were,
inevitably produce a close match of calculation with experiment.

You remark that my interpretation of the experiments in the Puzzles must be
wrong.   Of course it would be a rash scientist who thought (s)he could
never be wrong, so indeed I might be !   There are many interesting aspects
to the Puzzles, to my mind.   One is that the original workers offered no
explanation, and, indeed, did not even notice, in many Puzzles, that there
was a problem !   To illustrate that, I cite Puzzles 2, 3, 4, 5 (especially)
and 9.

Another interesting aspect of the Puzzles is the very wide range of
phenomena to which they relate.  Some do not very obviously fall within the
ambit of "Physics" as such.  Puzzle 3 is an exercise in the classical
chemistry of spread films.

A further interesting aspect of the Puzzles is that the true side-by-side
model, depending on your point of view!, offers a coherent and unifying
explanation of each and every one of the Puzzles in an unforced way.

The Puzzles recall the experimental results of research teams and often
compare them, one with another. If the double helix is the true structure of
DNA, are we going to need about a dozen different explanations of the 19 or
20 Puzzles, case by case, when the original workers could not explain their
results, or did not address the contradictions ?   Or is there just nothing
to explain ?

As you say, solid fibres of DNA are not actually solid, and, being drawn
from concentrated aqueous solutions, they do contain water.   In Puzzle 6,
to form a triple helix, as far as I can understand what is necessary, one
duplex must unwind, albeit slowly, and another duplex, presumably adjacent,
must rotate, anyway locally, if not along its whole length simultaneously.
I assume that that process requires torque somewhere, and, because it would
take time, there would be some kind of angular velocity.  You are right that
angular velocity would be quickly dampened by frictional forces.  That is
why I don't think the process can occur at all.  I deduce that the change of
duplex to triplex takes several days because that time is needed by the
bases, alone, to rotate to find new partners in a side-by-side triplex.

And then there is the problem of the random axial translation of the ends of
the duplexes.  It could be possible for the same duplex to be unwinding at
one end, and forming a triplex at the other, according to the plectonaemic
model.

One interesting aspect of these dynamics, whatever they actually do consist
of, is that the torque in any plectonaemic model of duplex changing to
triplex, wherever it arises, is a vector, whereas a move to a lower energy
state involves energy dissipation in all directions, i.e., a scalar.  Can a
vector be invoked to effect a change in a scalar in such a situation ?

You are right that I would not understand well enough the physics of the
rotational motion of large suspended masses (presuming that the changing
rotation of the Earth on its elliptical orbit, say, and other factors, were
properly discounted also !).  But then, even today, physicists do not always
get it right !  What about cold fusion, for example ?

I shall press on to post revised versions of the Puzzles, with the true
side-by-side solution.   Apart from being told that I'm wrong, nobody has
offered another solution to the Puzzles overall, nor explained why there
isn't a problem !

Clive Delmonte
-----------------------------------------------------------



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M________\ #_________________________________________________
;__________________________________ `
`
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