DNA Structure: Puzzle #7 (Version 2)

Clive Delmonte clived at ndirect.co.uk
Wed Mar 31 11:08:12 EST 1999


The Unwinding Problem Inside Fibres

Leslie et al.(19) reported that a fibre of poly(dI).poly(dC), in the absence
of unwinding proteins, which gave sharp X-ray diffraction spots shortly
after being drawn, was unstable and transformed irreversibly after a few
days into poly(dC).poly(dI).poly(dC+). This new, three-stranded molecule
also gave sharp spots in its own X-ray diffraction pattern which did not
have any of the original spots.

Therefore the conversion was complete. Very similar results had been
reported for fibres of poly(dA).poly(dT) converting to
poly(dT).poly(dA).poly(dT) (20) and for the formation of
poly(U).poly(A).poly(U) from poly(U).poly(A) (21).

The problem is evident:  How and where would the torque arise, and be
sustained, inside a solid fibre that would permit one double helix to rotate
so as to unwind one strand and rewind it onto an adjacent triple helix
having a different diameter and rotating at a different angular velocity,
the conversion being complete ?

Inside the fibre, individual molecules would have a random axial translation
so only very rarely would two adjacent double helices have the same axial
starting point down the fibre, and they would be unlikely to be perfectly
straight inside the fibre but would probably be entwined around neighbours.

A true side-by-side duplex DNA structure (see attachment) could form a
side-by-side triplex, without any unwinding, by the linear transfer of a
strand across to a neighbouring duplex.
-----------------------------------------------------------------
19        Polymorphism of DNA Double Helices; A.G.W. Leslie et al.; J Mol
Biol Vol 143 (1980) 49 - 72

20      Structures for the Polynucleotide Complexes Poly(dA).Poly(dT) &
Poly(dT).Poly(dA).Poly(dT); Struther Arnott & E. Selsing; J Mol Biol Vol 88
(1974) 509-521

21      Structures for Poly(U).poly(A).poly(U) Triple Stranded
Polynucleotides; Struther Arnott & P.J. Bond; Nature New Biology Vol 244
(1973) 99-101

------------------------------------------------------------------
Clive Delmonte









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M`'____________________________________________\/X/__________
M_______P`/__________W\__P !_________________________________
M____________#^#_________________\ #__________- at __]__?_______
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M__]P'/_________________P`/_________\?_______________________
M____________________________?WS_________________\ #_________
M_'___________________________________________________W]\____
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M____________________________________________``#_____________
M____\ #_____________________________________________________
M_________P``__________________ `____________________________
M__________________________________]__/_________________P`/__
M____________________________________________________________
M?_S_________________\ #_____________________________________
M_________________________W_\__________________ `____________
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M____________________________________\ #_____________________
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M__ `________________________________________________________
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M__________________ `________________________________________
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M___P`/______________________________________________________
M____________________________\ #_____________________________
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M___________________P`/______________________________________
M____________________________________________\ #_____________
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M__________________________ `________________________________
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M________________\ #_________________________________________
M__________________________________________ `________________
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M_______P`/__________________________________________________
M________________________________\ #_________________________
M__________________________________________________________ `
M____________________________________________________________
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M________________________________________________\ #_________
M____________________________________________________________
M______________ `____________________________________________
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M____\ #_____________________________________________________
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M____________________\ #_____________________________________
M______________________________________________ `____________
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M___P`/______________________________________________________
M____________________________\ #_____________________________
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M`/__________________________________________________________
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M____________\ #_____________________________________________
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M___P`/______________________________________________________
M____________________________\ #_____________________________
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M________________\ #_________________________________________
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M____________________________________________________________
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M____________________________________________________________
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M____\ #_____________________________________________________
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M____________________________________________________________
M________\ #_________________________________________________
;__________________________________ `
`
end





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