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ENDLESS FORMS: SPECIES AND SPECIATION
Symposium in Honor of Guy L. Bush
Organizers: Daniel Howard and Stewart Berlocher
On May 19-May 23, 1996, a symposium will be held at
Asilomar, California to summarize our current understanding
of speciation and to plot directions for future research.
Since the last symposium on speciation in 1987, which
resulted in the highly regarded "Speciation and its
Consequences" (D. Otte and J. Endler, eds.), significant
progress has been made in a number of areas, including
genetic change at speciation, the molecular genetics of
reproductive isolation, models of speciation, and
speciation in the fossil record. Controversy over species
concepts, ever simmering, is close to the boiling point at
present.
The symposium will honor Guy Bush on the occasion of the
30th anniversary of his 1966 Museum of Comparative Zoology
monograph on *Rhagoletis*; this watershed work established
these fruit flies as a touchstone of debate about the
possibility of sympatric speciation via ecological shifts.
Bush has long been associated with challenging accepted
ideas on speciation, and wrote a key review paper in 1975
that set the stage for modern speciation research.
Although the symposium will honor Bush, its major purpose
is to bring some of the world's leading researchers on
speciation together in a gathering structured to encourage
the free exchange of ideas and data. Expanded versions of
talks by the 30 participants will be published in a
symposium volume of the same title as the symposium.
MAJOR AREAS TO BE COVERED:
* Species Concepts
* Geography, Ecology, and Population Structure
* The Nature of Mate Recognition and Reproductive Isolation
* The Genetics of Reproductive Isolation
* Interactions Between Species and the Nature of Species
Boundaries
* The Ecology of Speciation and the Evolution of Novelty
SPEAKERS
Michael Arnold Mark MacNair James Patton
Stewart Berlocher Jim Mallet Michael Ritchie
Guy Bush Therese Markow Bill Rice
Roger Butlin Marta Martinez Wells Louise Roth
Jeff Feder Amy McCune Dolph Schluter
Rosemary Grant Steph Menken Kerry Shaw
Richard Harrison John Mercer Franco Spirito
Daniel Howard Horacio Naviera Alan Templeton
Paul Johnson Stephen Palumbi David Wake
Haris Lessios Dorothy Pashley Jack Werren
TALKS:
*SPECIES CONCEPTS*
KERRY SHAW
The diversity of process and the evolution of natural
groups
ALAN TEMPLETON
Geography, population structure, ecology, and gene trees.
*GEOGRAPHY, ECOLOGY, AND POPULATION STRUCTURE*
STEWART H. BERLOCHER
The biogeographic/phylogenetic evidence for sympatric
speciation.
PAUL JOHNSON
Multi-locus models of sympatric speciation.
JEFFREY L. FEDER
Host race formation and sympatric speciation in the apple
maggot fly: Fact or fiction?
WILLIAM R. RICE
The evolution of reproductive isolation without allopatry:
the role of sexually antagonistic coevolution.
STEPHANUS B. J. MENKEN
Evolution of insect-plant associations: *Yponomeuta* as a
case study.
AMY MCCUNE
The mire of speciating fishes in ancient lakes: Could
speciation be sympatric?
DOLPH SCHLUTER
Ecological causes of speciation.
M. R. MCNAIR
The evolution of edaphic endemics.
HARIS A. LESSIOS
The first stages of speciation as seen in organisms
separated by the Isthmus of Panama.
DAVID B. WAKE
Genetic interactions in a ring species: Rings within
rings, and weak links.
JAMES L. PATTON
Ridges, rivers, and refuges: The timing and
diversification of Amazonian Mammals.
*THE NATURE OF MATE RECOGNITION AND REPRODUCTIVE ISOLATION*
THERESA ANN MARKOW
Reproductive isolation in *Drosophila* : A case study from
the Sonoran desert.
MARTA L. MARTINEZ WELLS
The role of mating signals in reproductive isolation among
cryptic species of insects.
STEPHEN R. PALUMBI
Speciation and gamete recognition: Patterns of
polymorphism of the gamete recognition protein bindin in
sea urchins.
DANIEL J. HOWARD
The evolution of barriers to fertilization in terrestrial
organisms.
JOHN H. WERREN
Symbiotic bacteria and speciation.
*GENETICS OF REPRODUCTIVE ISOLATION AND MATE RECOGNITION*
HORACIO NAVIERA
The genetics of hybrid male sterility in *Drosophila*.
MICHAEL G. RITCHIE
The evolutionary genetics of sexual signaling: Is there an
equivalent of Haldane's Rule for premating isolation?
DOROTHY PROWELL PASHLEY
Linkage, sex, and speciation in Lepidoptera.
FRANCO SPIRITO
The role of chromosomal change in speciation.
JAMES MALLET
Mimicry and warning color at the boundary between
microevolution and macroevolution.
*INTERACTIONS BETWEEN SPECIES AND THE NATURE OF SPECIES
BOUNDARIES*
RICHARD G. HARRISON
Molecular genetic markers and the study of speciation.
MICHAEL L. ARNOLD
Assortative mating and selection in a hybrid population of
*Iris*.
ROGER BUTLIN
What do hybrid zones in general, and the *Chorthippus
parallelus* zone in particular, tell us about speciation?
B. ROSEMARY GRANT
Hybridization and speciation in Darwin's Finches.
*THE EVOLUTION OF NOVELTY*
V. LOUISE ROTH and JOHN M. MERCER
Bushy radiations and diversification in the Sciuridae.
*CONCLUSION*
GUY L. BUSH
Historical perspective and conclusions.
MEETING FORMAT AND SCHEDULE:
In the style of Gordon Conferences, there will be no
concurrent sessions, and much free time for discussion.
Arrivals will be Sunday May 19 and departures will be
Thursday May 23. During the three full days of the meeting
(Monday May 20 through Wednesday May 22) talks are
scheduled for morning and early afternoon, leaving late
afternoons and evenings free. Each day immediately
following the last afternoon session there will be a one
hour discussion of the day's talks, with the speakers
present. A banquet will conclude the meeting.
POSTER SESSION
In order to allow as many attendees as possible to
participate and contribute ideas, space for 60 posters will
be available in a room adjacent to the lecture room.
ADDITIONAL INFORMATION
Background information will be available soon on the World
Wide Web at http://www.life.uiuc.edu/berlocher/symposium/
REGISTRATION INFORMATION:
Registration for the meeting will be handled by the
organizers, while room and board at Asilomar will be
handled directly by Asilomar. To obtain all registration
information, please contact Daniel Howard at New Mexico
State via email or conventional mail. We will return the
meeting and Asilomar forms to you.
ORGANIZERS
Daniel Howard
Department of Biology
New Mexico State University
Las Cruces, NM 88003
dahoward at nmsu.edu
Stewart Berlocher
Department of Entomology/320 Morril Hall
University of Illinois
505 S. Goodwin
Urbana, IL 61801
stewart_berlocher at qms1.life.uiuc.edu
IMPORTANT NOTE
*THE MEETING WILL LIMITED TO THE FIRST 200 REGISTRANTS*
REGISTRATION DEADLINE:
All registration must be completed by January 1, 1996
in order to make preparations with Asilomar.
ACCOMMODATIONS AT ASILOMAR AND ELSEWHERE:
Lodging at Asilomar is comparable in price with local
accommodations (see below), is attractively located on the
coastline, and is, of course, very convenient to the
meeting and attendant discussion. However, participants
need not stay at Asilomar to attend, and those traveling
with spouses/families may want to investigate staying in
nearby Monterey.
MEETING COSTS:
MEETING REGISTRATION -
The cost of meeting registration is commensurate with
that of meetings such as the Society for the Study of
Evolution. Registration is $140, and $110 for
students. (Finances will be handled by New Mexico
State at Las Cruces. Funds will be used exclusively
for expenses associated with the meeting, such as
travel expenses for speakers and rental of audio-visual
equipment. If money remains after meeting expenses
have been covered, it will be refunded to
participants).
ACCOMMODATION AT ASILOMAR -
Current prices for a 4 day meeting range between $477
and $249 per person, depending on accommodations and
occupants/room. PLEASE NOTE THAT THIS IS INCLUDES BOTH
ROOM AND BOARD (food at Asilomar is reputedly quite
good), FOR THE ENTIRE DURATION OF THE MEETING.
Asilomar has set aside accommodations for 238 persons,
so some spouses/ families can be accommodated at
Asilomar. As with meeting registration, accommodations
at Asilomar are on a "first come-first served" basis.
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Digital *Rhagoletis pomonella*, the apple maggot fly
